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Zanthoxylum kauaense

A.Gray (1854)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Zanthoxylum

a‘e [ae], aʻe [ae], hea‘e, mānele, zanthoxylum

Description

Key Characters:

Growth Form: Small to medium-sized trees (1–)3–15 m tall.

Stems: Trunk to 40–50 cm in diameter, bark brown to grayish brown, new growth glabrous or finely puberulent, young branches subtended by a pair of bud scales.

Roots:

Leaves: Leaves compound. Alternate. Leaflets ovate to elliptic, sometimes broadly elliptic, broadly elliptic–ovate, or suborbicular, rarely lanceolate, lateral ones usually asymmetrical; leaflets 3–7(–11), thick,4–19(–21) cm long, 2–10(–14) cm wide, lateral petiolules articulated only at base, 0–7(–10) mm long, terminal petiolule once or twice articulated, at base and ± near leaflet, 10–75 mm long. Apex obtuse to rounded, sometimes bluntly acute, rarely acute or weakly acuminate. Base cuneate to rounded, subtruncate, or subcordate, adaxial side ± terminating higher. Surfaces glabrous or sometimes finely puberulent, and then more densely so when young, glandular punctate; Blades coriaceous. Margins entire or sometimes minutely crenulate. Lateral veins 5–10 pairs, spreading, higher order venation forming a conspicuous reticulate pattern. Rachis terete (cylindrical) or winged. Stipules absent.

Flowers: Flowers usually 15–150 or more in axillary, open, cymose inflorescences usually 8–15 cm long, glabrous or finely puberulent, arising at beginning of new growth on stem, and stem continuing vegetatively, peduncles 10–70 mm long, pedicels 1–5 mm long, subtended by a pair of bud scales; bracteoles ca. 1 mm long, usually caducous. Flowers unisexual, actinomorphic. Calyx of 4–5 sepals; sepals deltate, 0.5–1 mm long, ± finely puberulent, ciliolate. Corolla of 4–5 petals, white, sometimes pink or red-tinged, oblong–ovate to oblong–elliptic or oblong–lanceolate, 4–5 mm long. Stamens (3)4–5, alternate with the petals, glabrous, absent in pistillate flowers; anthers medifixed, dithecal. Ovary superior, 1-carpellate; ovule placentation axile, rudimentary in staminate flowers; style oblique; stigma capitate.

Fruit: Follicles 1; falcate; obovoid; 8–12(–15) mm long; apex rounded; exocarp rugulose and pitted. Seeds 1 per fruit; on a slender funicle; black; subglobose; compressed; 7–13 mm in diameter.

Ploidy:

Habitat: Mesic forest; sometimes in dry to wet forest.

Elevation Range: 300–1,980 m.

Historical Distribution

Images

Synonyms (49)

Fagara glandulosa (Hillebr.) Engl. (1896)
Fagara glandulosa var. hiloensis Sherff (1958)
Fagara kauaense (A.Gray) Engl. (1896)
Fagara kauaense var. kohalana Sherff (1958)
Fagara kauaense var. kohuana Skottsb. (1944)
Fagara kauaense var. tenuifolia O.Deg. & Sherff (1958)
Fagara kauaensis (A.Gray) Engl. (1896)
Fagara kauaensis var. kohalana Sherff (1958)
Fagara kauaensis var. tenuifolia O.Deg. & Sherff (1958)
Fagara kauaiensis var. tenuifolia Deg. & Sherff
Fagara mauiensis (H.Mann) Engl. (1896)
Fagara mauiensis f. glabra (Rock) Skottsb.
Fagara mauiensis var. anceps (Rock) Skottsb. (1958)
Fagara mauiensis var. cranwelliae Skottsb. (1944)
Fagara mauiensis var. kaalana Sherff (1958)
Fagara mauiensis var. lanaiensis Sherff (1958)
Fagara mauiensis var. maunahuiensis Sherff (1958)
Fagara mauiensis var. rigida (Rock) Sherff (1958)
Fagara maviensis
Fagara maviensis f. petiolulata (Rock) H.St.John (1945)
Fagara maviensis var. anceps (Rock) H.St.John (1945)
Fagara maviensis var. cranwelliae
Fagara multiflora Engl. (1896)
Fagara multifoliolata O.Deg. & Skottsb. (1936)
Fagara semiarticulata (H.St.John & Hosaka) O.Deg. (1936)
Fagara semiarticulata f. laiense O.Deg. (1936)
Fagara semiarticulata var. sessilis O.Deg. (1936)
Fagara skottsbergii O.Deg. & I.Deg. (1960)
Zanthoxylum glandulosum Hillebr. (1888)
Zanthoxylum hillebrandii P.G.Waterman (1975)
Zanthoxylum hillebrandii var. hiloense (Sherff) H.St.John (1976)
Zanthoxylum kauaense var. kohalanum (Sherff) H.St.John (1973)
Zanthoxylum kauaense var. kohuanum (Skottsb.) H.St.John (1973)
Zanthoxylum kauaense var. tenuifolium (O.Deg. & Sherff) H.St.John (1976)
Zanthoxylum maviense H.Mann (1866)
Zanthoxylum maviense f. glabrum Rock (1913)
Zanthoxylum maviense f. petiolulatum Rock (1913)
Zanthoxylum maviense var. anceps Rock (1913)
Zanthoxylum maviense var. cranwelliae (Skottsb.) H.St.John (1973)
Zanthoxylum maviense var. kaalanum (Sherff) H.St.John (1976)
Zanthoxylum maviense var. lanaiense (Sherff) H.St.John (1976)
Zanthoxylum maviense var. maunahuiense (Sherff) H.St.John (1976)
Zanthoxylum maviense var. rigidum Rock (1913)
Zanthoxylum molokaiense B.C.Stone (1985)
Zanthoxylum semiarticulatum H.St.John & Hosaka (1932)
Zanthoxylum semiarticulatum f. laiense (O.Deg.) Fosberg (1969)
Zanthoxylum semiarticulatum var. sessile Fosberg (1969)
Zanthoxylum semiarticulatum var. sessilis (O.Deg.) Fosberg (1969)
Zanthoxylum skottsbergii (O.Deg. & I.Deg.) H.St.John (1973)

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: U.S. Expl. Exped., Phan. 1: 354 (1854)

Other References

Wagner et al. 1990:1215 (K, O, Mo, L, M, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Zanthoxylum kauaense West Maui, Lahainaluna Preserved_Specimen Hillebrand, W.B. Maui BISH
2 Zanthoxylum kauaense Bird Park Kipuka puaulu moderate forest Preserved_Specimen Morley, H. Hawaii BISH 1934-08-11
3 Zanthoxylum kauaense Pupukea-Kahuku Trail Preserved_Specimen Krauss, N.H. Oahu BISH 1933-12-17
4 Zanthoxylum kauaense Waianae Mts., Kahanakaiki Valley, Piko Trail Preserved_Specimen Krauss, N.H. Oahu BISH 1933-12-30
5 Zanthoxylum kauaense Lanai Preserved_Specimen Hillebrand, W.B. Lanai BISH
6 Zanthoxylum kauaense Waimea Preserved_Specimen Hillebrand, W.B. Kauai BISH
7 Zanthoxylum kauaense Waianae Mts., near Halona Valley Preserved_Specimen Welch, D.A. Oahu BISH 1936-06-04
8 Zanthoxylum kauaense Lihue E. side Preserved_Specimen Russ, G.W. Kauai BISH 1929-10-01
9 Zanthoxylum kauaense S.slope of Hauula Ridge, 10 ft. below trail, (Kaipapau Forest Reserve) Preserved_Specimen Suehiro, A. Oahu BISH 1933-10-15
10 Zanthoxylum kauaense West Maui, Maunahuoma Preserved_Specimen Forbes, C.N. Maui BISH 1910-05-01
11 Zanthoxylum kauaense Kawaihae uka, Kohala Preserved_Specimen Hillebrand, W.B. Hawaii BISH 1862-01-01
12 Zanthoxylum kauaense Makua Preserved_Specimen Russ, G.W. Oahu BISH 1930-06-01
13 Zanthoxylum kauaense Kanehoa, East side Preserved_Specimen Russ, G.W. Oahu BISH 1930-06-19
14 Zanthoxylum kauaense Kepuka Puaulu. Preserved_Specimen Russ, G.W. Hawaii BISH 1927-08-01
15 Zanthoxylum kauaense Waianae Mts., Makaha Valley Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
16 Zanthoxylum kauaense Makaleha Preserved_Specimen Russ, G.W. Oahu BISH 1929-10-01
17 Zanthoxylum kauaense Palikea Preserved_Specimen Swezey, O.H. Oahu BISH 1935-02-03
18 Zanthoxylum kauaense Back of Kilauea Preserved_Specimen Lydgate, J.M. Hawaii BISH 1909-07-01
19 Zanthoxylum kauaense Kapapala Preserved_Specimen Russ, G.W. Hawaii BISH 1928-03-01
20 Zanthoxylum kauaense Maunakapu Preserved_Specimen Suehiro, A. Oahu BISH 1931-11-22
21 Zanthoxylum kauaense East Lanai, Awehi, 4 gulches over S of Puhielelelu Ridge Preserved_Specimen Montgomery, S.L. Lanai BISH 1971-06-06
22 Zanthoxylum kauaense East Lanai, Awehi Preserved_Specimen Montgomery, S.L. Lanai BISH 1971-06-09
23 Zanthoxylum kauaense Kawaihae uka Preserved_Specimen Hillebrand, W.B. Hawaii BISH
24 Zanthoxylum kauaense Waianae Mts, Puu Kaua Preserved_Specimen O'Connor, P.J. Oahu BISH 1982-10-10
25 Zanthoxylum kauaense Hilo Preserved_Specimen Lydgate, J.M. Hawaii BISH
26 Zanthoxylum kauaense Wahiawa Mts Preserved_Specimen Lydgate, J.M. Kauai BISH
27 Zanthoxylum kauaense Polihua, trail to Preserved_Specimen Judd Jr., A.F. Lanai BISH 1913-08-01
28 Zanthoxylum kauaense Waimea Preserved_Specimen Mann, H. Kauai BISH 1864-01-01
29 Zanthoxylum kauaense Kaaikuhae Preserved_Specimen Russ, G.W. Oahu BISH 1929-06-01
30 Zanthoxylum kauaense Kaala Mt Preserved_Specimen Hillebrand, W.B. Oahu BISH 1861-06-01
31 Zanthoxylum kauaense Kipuka Puaulu Preserved_Specimen Russ, G.W. Hawaii BISH 1930-06-01
32 Zanthoxylum kauaense Pupukea-Kahuku Trail, Koolau Mts Preserved_Specimen Furmidge, G. Oahu BISH 1933-02-19
33 Zanthoxylum kauaense Waiapaa Preserved_Specimen Munro, G.C. Lanai BISH 1915-03-18
34 Zanthoxylum kauaense Koolau Mountain Range: Makaha Valley Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
35 Zanthoxylum kauaense Waianae Mts., Makaha Valley Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
36 Zanthoxylum kauaense Awehi, E Lanai Preserved_Specimen Montgomery, S.L. Lanai BISH 1971-06-06
37 Zanthoxylum kauaense Lanai Preserved_Specimen Hillebrand, W.B. Lanai BISH
38 Zanthoxylum kauaense Waianae Mts., Makaha Valley Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
39 Zanthoxylum kauaense W Maui Preserved_Specimen Maui BISH
40 Zanthoxylum kauaense Maunahui Preserved_Specimen Hillebrand, W.B. Molokai BISH
41 Zanthoxylum kauaense [Hawaiian Islands; U.S.A.; Molokai; Hawaii; Maui Preserved_Specimen Hillebrand, W.B. Molokai BISH
42 Zanthoxylum kauaense Lahainaluna Preserved_Specimen Hillebrand, W.B. Maui BISH
43 Zanthoxylum kauaense Kula, S of sanitorium Preserved_Specimen Forbes, C.N. 2176 Maui BISH 1920-04-08
44 Zanthoxylum kauaense Haleakala, Makawao, Waihou Gulch Preserved_Specimen Rock, J.F.C. 10200 Maui BISH 1912-03-01
45 Zanthoxylum kauaense Puaulu, Kilaueu Preserved_Specimen Rock, J.F.C. 10202 Hawaii BISH 1911-07-01
46 Zanthoxylum kauaense trail to Waialae Valley Preserved_Specimen Rock, J.F.C. 5960 Kauai BISH 1909-09-06
47 Zanthoxylum kauaense Puuwaawaa Preserved_Specimen Forbes, C.N. 61 Hawaii BISH 1911-06-08
48 Zanthoxylum kauaense Near Waiakeakua Preserved_Specimen Munro, G.C. 423 Lanai BISH 1915-04-23
49 Zanthoxylum kauaense Kaiholena Valley Preserved_Specimen Forbes, C.N. 2 Lanai BISH 1913-06-01
50 Zanthoxylum kauaense ridge behind Papaa, Moloaa FR Preserved_Specimen Degener, O. 28096 Kauai BISH 1952-01-16
51 Zanthoxylum kauaense Pupukea Military Trail Preserved_Specimen MacDaniels, L.H. 544 Oahu BISH 1927-01-29
52 Zanthoxylum kauaense West Mau/Molokai Preserved_Specimen Hillebrand, W.B. BISH
53 Zanthoxylum kauaense Limahuli Valley wet pali; ohia, Eugenia, uluhe; 80 deg. slope Preserved_Specimen 30' x 10 tree Perlman, S. 189 Kauai BISH 1978-07-05
54 Zanthoxylum kauaense ridge crest between Kaua and Kanehoa Chamaesyce clearing Preserved_Specimen small Takeuchi, W.N. 2847 Oahu BISH 1986-09-20
55 Zanthoxylum kauaense Near summit of Pupukea-Kahuku trail Exposed rain-forest on ridge Preserved_Specimen Degener, O. 8600 Oahu BISH 1929-03-31
56 Zanthoxylum kauaense Waianae Mts., trail on E ridge of Puu Kaua N facing slope Preserved_Specimen 4 m tree; many fruit, ~18/infructesence on branches Marr, K.L. 24 Oahu BISH 1988-04-17
57 Zanthoxylum kauaense near head of Kaluaa base mesic forest Preserved_Specimen 4-5 m tree Takeuchi, W.N. 3710 Oahu BISH 1987-11-08
58 Zanthoxylum kauaense Waimea Canyon, toward head from Puu Kapele dryland forest Preserved_Specimen 12' x 5 tree; inodorous Christensen, C. 199 Kauai BISH 1977-05-07
59 Zanthoxylum kauaense Honopu, drainage north of trail. Acacia koa-Metrosideros polymorpha montane mesic forest. W/ Zanthoxylum dipetalum, Dicranopteris linearis, hedyotis terminalis, Pleomele aurea, Bobea brevipes, Psychotria mariniana, Melicope anisata, M. barbigera, Pouteria sandwicensis, Dodonaea viscosa, Di Preserved_Specimen tree, 3.5 m tall, trunk mottled gray with brown blotching, with Zanthoxylum dipetalum, Claoxylon sandwicensis, Kadua affinis, Xylosma hawaiiensis, Cyrtandra longifolia Wood, K.R. 14006 Kauai BISH 2009-11-10
60 Zanthoxylum kauaense South Opaeula Gulch, Paalaa, Koolau Mts wooded ridge Preserved_Specimen small tree St.John, H. 10635 Oahu BISH 1930-11-09
61 Zanthoxylum kauaense Mt. Kaala, DuPont Trail, Mokuleia FR dry forest on ridge Preserved_Specimen 7 m x 2 dm tree St.John, H. 23693 Oahu BISH 1949-05-22
62 Zanthoxylum kauaense Waimea District, Nuololo, central branch Metrosideros-Acacia mesic forest with Alphitonia ponderosa, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bobea brevipes, Melicope ovata, M. barbigera Preserved_Specimen tree, 3.75 m tall, stems mottled dark brown, gray Wood, K.R. 15042 Kauai BISH 2012-06-20
63 Zanthoxylum kauaense Puu Kapele forest Preserved_Specimen Rock, J.F.C. Kauai BISH 1956-02-01
64 Zanthoxylum kauaense ridge S of S. Opaeula Gulch, Paalaa, Koolau Mts wooded Preserved_Specimen St.John, H. 11523 Oahu BISH 1932-01-17
65 Zanthoxylum kauaense Olaa Forest Reseve, behind Volcano dump Wet forest habitat. Preserved_Specimen 16m x 11.5cm Marr, K.L. 81 Hawaii BISH 1988-07-30
66 Zanthoxylum kauaense Waianae Mts., below Puu Kalena with Metrosideros, Myrsine lessertiana, Melicope, Diospyros, Freycinetia, Scaevola gaudichaudii, Dicranopteris, Labordia, Pipturus, Psidium Preserved_Specimen tree Perlman, S. 9404 Oahu BISH 1988-01-15
67 Zanthoxylum kauaense Makaha Valley, N of Puu Kawiwi, Kamaileunu Ridge, Waianae Mts crest of minor spur; with Schinus, Acacia, Diospyros Preserved_Specimen 15' 3 tree Nagata, K.M. 1101 Oahu BISH 1972-12-28
68 Zanthoxylum kauaense Awaawapuhi trail, Honopu rainforest Preserved_Specimen Woolford, H.A. 430 Kauai BISH 1956-12-29
69 Zanthoxylum kauaense Waimea District. Nuololo, southern branch. Metrosideros-Acacia montane mesic forest, 80% closed canopy with Alphitonia ponderosa, Antidesma platyphylla, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bobea brevipes, Melico Preserved_Specimen tree, 4 m tall, few branched, stems dark brown with lichen, with fruit, with Ilex anomala, 4 trees in area. Wood, K.R. 14926 Kauai BISH 2012-04-13
70 Zanthoxylum kauaense North Kona District: Puuwaawaa, Waiho. 1/2 mile NNW of Hibiscadelphus grove Hidden in mamane trees. Preserved_Specimen One tree. 6 inches in diameter, 20 feet high. Male flower in parts of 4. Corolla white inside, purplish outside. Leaves: light green, dull; veins red above. Fagerlund, G.O. 1016 Hawaii BISH 1945-03-24
71 Zanthoxylum kauaense E rim of Kalalau valley forest Preserved_Specimen small tree Degener, O. 21470 Kauai BISH 1951-12-27
72 Zanthoxylum kauaense Kokee. Upper Mahanaloa Valley, just before yellow gate and picnic table Mixed mesic forest w thick patches of Lantana in valley bottom. Acacia koa - Metrosideros polymorpha w/ Tetraplasandra kavaiensis, Melicope ovata, Alyxia stellata, Dodonaea viscosa, Alphitonia, Leptecophylla, Wikstroemia furcata, Xylosma crenatum, X. hawaiiense, Coprosma, Myrsine lanaiensis, Carex wahuensis, Hedyotis terminalis, Scaevola. Preserved_Specimen Small tree about 6 ft tall., Bark smooth dark brown w/ white. LVs adaxial blade medium green glossy, abaxial light green, mostly odd-pinnate, but some evenly pinnate lvs. Red color at petiole junctions. Fl buds white w/ green tint. Nectar disk yellow. Tangalin, N. 1841 Kauai BISH 2008-11-19
73 Zanthoxylum kauaense Waianae Mts., Pukaloa Gulch, above Schofield, off ridge trail to Puu Kalena, near Kumakalii wet forest, with Metrosideros, Alyxia, Canthium, Nestegis, Chamaesyce, Psidium, Antidesma, Pleomele, Claoxylon, Urera, Nothocestrum longiflorum, Embelia, Pisonia, Bobea, Psychotria, Scaevola, Melicope, Astelia, Labordia, Perrottetia Preserved_Specimen Perlman, S. 5644 Oahu BISH 1987-04-16
74 Zanthoxylum kauaense Kauai Hanalei District; Honopu Rim. Acacia koa-Metrosideros polymorpha-Dicranopteris linearis montane mesic forest w/ Alphitonia ponderosa, Melicope feddei, Antidesma platyphylla var. hillebrandii, Planchonella sandwicensis, Nestegis sandwicensis, Bobea brevipes, Melicope barbigera, Dodonaea viscosa, Kadua affinis Preserved_Specimen Tree, 5 m tall, stems mottle gray, brown, green, 2 trees in area, with fruits, mapped Wood, K.R. Kauai BISH 2015-07-10
75 Zanthoxylum kauaense Pupukea FR, Koolauloa Metrosideros forest Preserved_Specimen 18' tree Grant, M.L. 7157 Oahu BISH 1934-08-01
76 Zanthoxylum kauaense Olaa Forest Reseve, behind Volcano dump Other species: Myrsine, Psychotria, Pipturus, Cyrtandra, Gouldia, Cheirodendron, Freycinetia, Metrosideros, Cibotium. Preserved_Specimen Not healthy, leaves only from buds on larger branches. No infloresence. Leaflets 3-5, up to 7 on young stems. One sapling, 8 seedlings. 12m x 19cm DBH. Leaves with citrus odor, but not as strong as other individuals in this area. Marr, K.L. 76 Hawaii BISH 1988-07-29
77 Zanthoxylum kauaense near head of Kaluaa base mesic forest Preserved_Specimen 4-5 m tree Takeuchi, W.N. 3710 Oahu BISH 1987-11-08
78 Zanthoxylum kauaense Maunauna Valley, 3 mi S of Puu Hapapa, Waianae Mts. At head of Valley along the ridge trail in dry open forest. A single tree growing with Koa, Straussia, Coprosma, Passiflora edulis, Lantana and Eugenia sandwicensis, Metrosideros Preserved_Specimen A small tree about 15 ft high and 4in. in diam Pearsall, G.A. Oahu BISH 1958-10-06
79 Zanthoxylum kauaense ridge flute near Kumakalii native forest Preserved_Specimen Takeuchi, W.N. 2832 Oahu BISH 1986-09-01
80 Zanthoxylum kauaense Honouliuli contour mauka slopes; mesic forest Preserved_Specimen shade leaves detached at node; usually trifolate, but occasional 5-, 7- pinnate Takeuchi, W.N. 3579 Oahu BISH 1987-05-31
81 Zanthoxylum kauaense NW boundary of Kaluaa, near head of basin. field no. Waianae 491 quality stand of native material, including Zanthoxylum skottsbergii, Alectryon mahoe, Cyanea membranacea, Nothocestrum longifolium, Viola tracheliifolia, and numerous other spp. Preserved_Specimen basal diameter = 5 in., height 4m Takeuchi, W.N. 2821 Oahu BISH 1986-09-07
82 Zanthoxylum kauaense Waihou Spring, Olinda on slopes near gulch bottom, shade, ash forest Preserved_Specimen 15' tree, in bud and flower Hobdy, R.W. 795 Maui BISH 1980-04-20
83 Zanthoxylum kauaense central ridge of Makaleha Valley forest Preserved_Specimen Degener, O. 21273 Oahu BISH 1951-01-12
84 Zanthoxylum kauaense near Pohakea, at the divide in rock crevices; with Schinus Preserved_Specimen 3-4 m Takeuchi, W.N. Oahu BISH 1983-08-01
85 Zanthoxylum kauaense East Maui, Haleakala, near Idlewild gulch Preserved_Specimen Rock, J.F.C. 8532 Maui BISH 1910-09-01
86 Zanthoxylum kauaense below Palikea, Waianae Range rainforest Preserved_Specimen Degener, O. 8595 Oahu BISH 1932-10-23
87 Zanthoxylum kauaense Waianae Mts., Mokuleia, Makaleha Trail Metrosideros and Pelea forest, lateral ridge Preserved_Specimen One tree, 7 m. x 8 cm. St.John, H. 25284 Oahu BISH 1954-02-28
88 Zanthoxylum kauaense Waianae Mts., between Puu Hapapa and Puu Kanehoa, southern gulch open, N facing slope Preserved_Specimen 4 m x 10 cm; crushed leaves with strong odor; leaflets 3-5'; fruit Marr, K.L. 30 Oahu BISH 1988-05-14
89 Zanthoxylum kauaense Puu Kawiwi-Mt. Kaala ridge, Waianae Mts., Makaha-Waianae Kai wooded Preserved_Specimen 7 m x 2 dm tree St.John, H. 17599 Oahu BISH 1935-03-31
90 Zanthoxylum kauaense Awaawapuhi Valley, Honopu In mesic forest Preserved_Specimen Small tree, lvs. 5-7-leafleted. N.v. a'e Stone, B.C. 1573 Kauai BISH 1956-12-29
91 Zanthoxylum kauaense near head of Kaluaa base mesic forest Preserved_Specimen 4-5 m tree Takeuchi, W.N. 3710 Oahu BISH 1987-11-08
92 Zanthoxylum kauaense Pahole Gulch, Waialua, Waianae Mts. Near crest of southwest wall of gulch Dense forest of Metrosideros and Diospyros hillebrandii with understory of Alyxia, Lantana and Euphorbia hillebrandii. Preserved_Specimen Tree 25 ft. ht., 5 in. dbh. Leaflets 3, rarely 5. Nagata, K.M. 1548 Oahu BISH 1978-01-31
93 Zanthoxylum kauaense Kokee. Kawaikoa Stream Trail Above trail on S. side of stream, 70 m before 1/2 mile marker, Growing between two Zanthoxylum depetalum (KM 45, 46) Preserved_Specimen 7 m. tall x 13.5 cm. DBH. Sterile. Bumps on leaves apparently due to insects. Marr, K.L. 47 Kauai BISH 1988-07-15
94 Zanthoxylum kauaense East Maui, Haleakala, near Idlewild gulch Preserved_Specimen Rock, J.F.C. 8532 Maui BISH 1910-09-01
95 Zanthoxylum kauaense East side of Kaaawa Gulch (north slope of Kaala) Rain-forest Preserved_Specimen Degener, O. 11813 Oahu BISH 1937-04-11
96 Zanthoxylum kauaense Koele mountains Preserved_Specimen Forbes, C.N. 209 Lanai BISH 1913-06-01
97 Zanthoxylum kauaense Waimea District, Awaawapuhi, steep north facing slopes SE of Kainamanu. Metrosideros-Cheirodendron montane wet, summer mesic forest w/ Alphitonia, Planchonella, Claoxylon sandwicense, Xylosma hawaiiensis, Antidesma, Bobea brevipes, Myrsine lanaiensis, Dodonaea, Wilkstroemia furcata, Leptecophylla tameiameiae, Kadua affinis, Melicop Preserved_Specimen Tree, 5 m tall, with immature fruit, vigorous, trees occasional/rare in area, mapped. Wood, K.R. 16758 Kauai BISH 2016-03-24
98 Zanthoxylum kauaense Near summit of Pupukea-Kahuku trail Exposed rain-forest on ridge Preserved_Specimen Degener, O. 8596 Oahu BISH 1932-01-24
99 Zanthoxylum kauaense Kukui Trail, Waimea Canyon upper forest Preserved_Specimen Degener, O. 23961 Kauai BISH 1955-09-20
100 Zanthoxylum kauaense Waianae Mtns. Puu Hapapa Just below crest of ridge south of summit in forest dominated by Schinus and Metrosideros. Preserved_Specimen Sterile, single tree Marr, K.L. 48 Oahu BISH 1988-06-17
101 Zanthoxylum kauaense Above Mokuleia Trail east of Hunter's cabin area Old tree with little foliage with Psychotria; Coprosma; Myrsine; Psidium cattleianum. Preserved_Specimen Welton, P. 813 Oahu BISH 1992-02-20
102 Zanthoxylum kauaense Waianae Mts., trail on E ridge of Puu Kaua N facing slope Preserved_Specimen fruit; leaflets 5-13 Marr, K.L. 23 Oahu BISH 1988-04-17
103 Zanthoxylum kauaense Waimea District, Kawai'iki, upper drainages above twin falls of Koa'ie Canyon Metrosideros polymorpha mixed mesic forest, Gahnia beecheyi, Dianella sandwicensis, Dubautia laevigata, Hedyotis terminalis, Cheirodendron trigynum, Psychotria mariniana, Poa sandwicensis, Bidens cosmoides, Peperomia macraeana, Peperomia latifo Preserved_Specimen Tree, 4 m tall, moderately branched, no flower or fruit, petiolules longer than usual, rare but occasional, with Z. dipetalum Wood, K.R. 9117 Kauai BISH 2001-09-25
104 Zanthoxylum kauaense ridge S of S. Opaeula Gulch, Paalaa, Koolau Mts wooded Preserved_Specimen 18' x 5 tree St.John, H. 11110 Oahu BISH 1931-06-28
105 Zanthoxylum kauaense Waianae Mts., Waianae Kai, saddle between Mt. Kaala and Puu Kawiwi, ~100 m from power lines with Cibotium, Bidens, Alyxia, Psidium, koa, ohia, Cyanea, Joinvillea Preserved_Specimen 6 m x 11 cm; sterile; 3 leaflets Marr, K.L. 66 Oahu BISH 1988-07-04
106 Zanthoxylum kauaense Kokee, Kaluapuhi trail wet forest Preserved_Specimen 20' x 2.5 tree; smooth, pale bark; short branches, leaves tufted at ends; female; petals white, ovary green Herbst, D.R. 2028 Kauai BISH 1971-07-28
107 Zanthoxylum kauaense north Honouliuli mesic slopes Preserved_Specimen 4 m x 20 cm tree Takeuchi, W.N. 3755 Oahu BISH 1988-01-10
108 Zanthoxylum kauaense Waimea Canyon, mile 11.3 on Hwy. 550 forest; steep dry slope Preserved_Specimen 25' fallen tree; leaves dark glossy green above with yellow green midrib, below yellow green with prominant reticulate veination; corolla in bud purple; open petals yellow green with purple tinting; cut branch ends fragrant Flynn, T.W. 2197 Kauai BISH 1987-05-15
109 Zanthoxylum kauaense Ekahanui-Kaua among boulder rocks; in Pelea Preserved_Specimen 4 m tree Takeuchi, W.N. 3554 Oahu BISH 1987-05-16
110 Zanthoxylum kauaense near head of Kaluaa base mesic forest Preserved_Specimen 4-5 m tree Takeuchi, W.N. 3710 Oahu BISH 1987-11-08
111 Zanthoxylum kauaense Kanehoa in dense Planchonella canopy Preserved_Specimen Takeuchi, W.N. 3436 Oahu BISH 1987-02-22
112 Zanthoxylum kauaense Kanehoa ridge Metrosideros, Pelea, Psychotria Preserved_Specimen 15 ft. Pyle, L. 2029 Oahu BISH 1985-03-03
113 Zanthoxylum kauaense Waianae Range, south of Palikea Moderately dry slope near the summit ridge Preserved_Specimen Degener, O. 10159 Oahu BISH 1936-01-05
114 Zanthoxylum kauaense East slope of Puu Kaua in dense wet forest Preserved_Specimen 10 ft. slender tree Degener, O. 10319 Oahu BISH 1936-01-19
115 Zanthoxylum kauaense Honouliuli contour mauka slopes; mesic forest Preserved_Specimen shade leaves detached at node; usually trifolate, but occasional 5-, 7- pinnate Takeuchi, W.N. 3579 Oahu BISH 1987-05-31
116 Zanthoxylum kauaense Waianae Mts., Palikea, E of summit N facing slope; with Cryptomeria japonica Preserved_Specimen mature fruit Marr, K.L. 5 Oahu BISH 1988-02-13
117 Zanthoxylum kauaense N side of Mt. Kaala moist forest, steep slope; with Metrosideros, Dubautia, Sophora Preserved_Specimen 12' x 5 tree; flowers white Hatheway, W.H. 335 Oahu BISH 1950-09-09
118 Zanthoxylum kauaense Waianae Range, Honouliuli, Puu Kanehoa dry wooded ridge Preserved_Specimen 6 m tree Webster, G.L. 1630 Oahu BISH 1948-06-13
119 Zanthoxylum kauaense Below Kawaiiki Ridge along the S. fork of Twin Falls. Ten fet above gulch bottom. Mixed Mesic forest. Canopy of Acacia koa and Metrosideros polymorpha w/ Pittosporum, Psychotria, Coprosma, Cyanea leptostegia, Asplenium, Cheirodendron trigynum, Dianella sandwicensis, Poa sandwicensis, Perrottetia sandwicensis Preserved_Specimen 7 ft tall small upright tree. Lots of hainging green immature frt. Lvs dark green w/ yellow mottling. Occasional in area. Tangalin, N. 2352 Kauai BISH 2010-06-02
120 Zanthoxylum kauaense upper Awaawapuhi In wet forest dominated by uluhe, ohia, manono, pilo Preserved_Specimen 4 m emergent from uluhe thicket, scattered individuals. Similar to Oahu's skottsbergii but with foliage less variable. Takeuchi, W.N. 3689 Kauai BISH 1987-10-25
121 Zanthoxylum kauaense Pahole Gulch, Waialua, Waianae Mts near crest of steep spur, south seciton of gulch. Canopy of Grevillea and Metrosideros Preserved_Specimen Small tree 20 ft. ht., 3 in. dbh. Corolla 4. Leaflets 5, rarely 3. Flowers pungent. Nagata, K.M. 1541 Oahu BISH 1978-01-31
122 Zanthoxylum kauaense Waianae Range, south of Palikea Moderately dry slope near the summit ridge Preserved_Specimen Degener, O. 10159 Oahu BISH 1936-01-05
123 Zanthoxylum kauaense S. Opaeula Gulch, Paalaa, Koolau Range, S ridge wooded Preserved_Specimen 18' x 5 tree St.John, H. 11110 Oahu BISH 1931-01-28
124 Zanthoxylum kauaense E of Kalalau Lookout forest Preserved_Specimen small tree Degener, O. 20600 Kauai BISH 1950-01-31
125 Zanthoxylum kauaense South Opaeula Gulch, Paala, Koolau Mts., wooded south ridge Preserved_Specimen St.John, H. 11089 Oahu BISH 1931-05-03
126 Zanthoxylum kauaense near Pohakea with Pelea peduncularis Preserved_Specimen a small shrub Takeuchi, W.N. Oahu BISH 1985-01-01
127 Zanthoxylum kauaense near Kolekole in remnant patch of intact forest. with Alectryon, Pleomele forbesii, Phyllostegia hirsuta Preserved_Specimen ~6 m pole tree Takeuchi, W.N. 2814 Oahu BISH 1986-09-06
128 Zanthoxylum kauaense Koa Niu, Keauhou Mauka Remnant of a mesic forest, now with scattered trees and kikuya grass. Preserved_Specimen Male tree, 15 ft. tall, 3 in. DBH. Pale green foliage; smooth gray bark; yellow-green scented flowers, visited by many introduced honey bees. This tree in good condition. Herbst, D.R. 2138 Hawaii BISH 1971-09-16
129 Zanthoxylum kauaense Pupukea- Kahuku Trail, Koolau Mts wooded ridge Preserved_Specimen 15' tree St.John, H. 11568 Oahu BISH 1932-03-06
130 Zanthoxylum kauaense Hanalei Distric; Kalalau Rim; Kalahu side below first Kalalau lookout; Steep Diverse Montane Mesic forest with; Poa mannii, Lysimachia glutinosa, Zanthoxylum kauaense, Nototrichium sp. nov., Labordia helleri, Dryopteris unidentata, Metrosideros, Coprosma, Peucedanum, Vaccinium dentatum, Lipochaeta, Wilkesia. Preserved_Specimen 3 m tall tree; in early fruit; vigorous, single plant Wood, K.R. 1147 Kauai BISH 1991-08-15
131 Zanthoxylum kauaense middle Palawai Ridge forest Preserved_Specimen Degener, O. 10863 Oahu BISH
132 Zanthoxylum kauaense Huliwai mixed native exotic forest, steep dirt slopes; with Alyxia Preserved_Specimen few fruit Takeuchi, W.N. 2845 Oahu BISH 1986-09-20
133 Zanthoxylum kauaense Honouliuli contour mauka slopes; mesic forest Preserved_Specimen shade leaves detached at node; usually trifolate, but occasional 5-, 7- pinnate Takeuchi, W.N. 3579 Oahu BISH 1987-05-31
134 Zanthoxylum kauaense Kalalau Valley, rim forest Preserved_Specimen Degener, O. 28102 Kauai BISH 1961-03-14
135 Zanthoxylum kauaense Kukui Trail, Waimea Canyon upper forest Preserved_Specimen 15' tree Degener, O. 23960 Kauai BISH 1955-09-20
136 Zanthoxylum kauaense Waianae Mts., Palikea trail, SE of summit with Cryptomeria Preserved_Specimen 5 m x 7 cm; leaves with 3 leaflets, dark green above, lighter below; sterile Marr, K.L. 58 Oahu BISH 1988-06-26
137 Zanthoxylum kauaense Near summit of Pupukea-Kahuku trail Exposed rain-forest on ridge Preserved_Specimen Crushed leaves have lemon odor. Degener, O. 4109 Oahu BISH 1931-12-06
138 Zanthoxylum kauaense Waianae Range, near Puu Kalena on the Kolekole side On ridge top among Metrosideros and Dicranopteris, rather moist habitat Preserved_Specimen One female tree, petals 4 Obata, J.K. 350 Oahu BISH 1978-03-05
139 Zanthoxylum kauaense Honouliuli contour mauka slopes; mesic forest Preserved_Specimen shade leaves detached at node; usually trifolate, but occasional 5-, 7- pinnate Takeuchi, W.N. 3579 Oahu BISH 1987-05-31
140 Zanthoxylum kauaense Hanalei District, Honopu Rim Acacia koa-Metrosideros polymorpha-Dicranopteris linearis montane mesic with Alphitonia ponderosa, Melicope feddei, Bobea brevipes, Melicope barbigera, Dodonaea viscosa, Kadua affinis, Doodia kunthiana Preserved_Specimen Tree, 4m tall, trunk gray, 8 cm diameter, single tree Wood, K.R. 15131 Kauai BISH 2012-08-02
141 Zanthoxylum kauaense Between Mauna Kapu and Palikea cloudy forest Preserved_Specimen dying tree Degener, O. 21028 Oahu BISH 1950-10-29
142 Zanthoxylum kauaense Mauna Kapu, Palikea, Waianae Range ridge top Preserved_Specimen Hutchison, P.C. 2818 Oahu BISH 1965-05-16
143 Zanthoxylum kauaense Kamoku, Kaiholena Gulch, S slope native scrub forest Preserved_Specimen 2 m shrubby tree, sparingly branched Hobdy, R.W. 2200 Lanai BISH 1984-10-22
144 Zanthoxylum kauaense Waianae Range, DuPont trail above Wailua wet forest Preserved_Specimen ~12' tree; leaves pungent, spicey when bruised Stone, B.C. 3182 Oahu BISH 1959-12-20
145 Zanthoxylum kauaense Makaha Valley, 1.5 mi down Ridge Road with Metrosideros, Acacia, Zanthoxylum dipetalum Preserved_Specimen 20' tree. Immature fruit covered with oil gland. Petioles of leaflets tinged reddish brown Flynn, T.W. 1218 Kauai BISH 1985-09-10
146 Zanthoxylum kauaense Waianae Mts., trail to Puu Kanehoa just below top of ridge on N facing slope Preserved_Specimen 5 m x 5.5 cm tree; leaflets 5-7; few fruit Marr, K.L. 38 Oahu BISH 1988-06-07
147 Zanthoxylum kauaense Keauhou Mauka, Koa Nui; In remnants of a mesic, summer-dry forest Preserved_Specimen male tree, 15 ft. tall; 3 DBH; foliage pale green; bark smooth, gray; flowers scented, yellow-green, being worked over by many introduced honey bees. Herbst, D.R. 2138 Hawaii BISH 1971-09-16
148 Zanthoxylum kauaense central ridge of Makaleha Valley forest near summit Preserved_Specimen Degener, O. 20759 Oahu BISH 1950-08-19
149 Zanthoxylum kauaense Haupu, talus slope, foot of cliff talus slope Preserved_Specimen Small tree. Ht. 2 m. MacDaniels, L.H. 882 Kauai BISH 1927-02-26
150 Zanthoxylum kauaense Kohua Ridge north facing slopes above Awini Stream Metrosideros-Acacia montane mesic forest with Alphitonia pondersoa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena aurea, Polyscias kavaiensis, Planch Preserved_Specimen tree, 4.5 m tall, trunk tan-brown, mottled, in flower, single tree Wood, K.R. 14937 Kauai BISH 2012-04-25
151 Zanthoxylum kauaense Waianae Range, DuPont trail above Wailua steep slope, wind swept wet forest Preserved_Specimen 18' tree; fruit green Stone, B.C. 3184 Oahu BISH 1959-12-20
152 Zanthoxylum kauaense Kokee stream forest Preserved_Specimen ~9-12' tree Degener, O. 21484 Kauai BISH 1951-12-31
153 Zanthoxylum kauaense Waianae Mts.; 1 of S gulches between Puu Hapapa and Kanehoa open; N facing slope; 40 m above bottom Preserved_Specimen 4 m x 5 cm; leaflets 3-7, strong odor when crushed Marr, K.L. 29 Oahu BISH 1988-05-04
154 Zanthoxylum kauaense Waianae Mts., Puu Kanehoa in moist forest, on trail Preserved_Specimen Tree 7 m. high; lvs.thick, coriaceous, 1st veins white; fr.dark green, with paler oil dots. Cowan, R.S. 1039 Oahu BISH 1948-06-13
155 Zanthoxylum kauaense Between Puu Manawahua and Mauna Kapu, Nanakuli side forest Preserved_Specimen Degener, O. 20847 Oahu BISH 1950-06-28
156 Zanthoxylum kauaense beginning of trail into Waikolu Valley from Waikolu Lookout in open Metrosideros polymorpha forest at edge of pali, with mats of Dicranopteris linearis and such shrubs as Pipturus rockii, Coprosma stephanocarpa, Lysimachia hillebrandii, Straussia, Dubautia scabra leiophylla Preserved_Specimen Shrub to 8 feet Carlquist, S. 2224 Molokai BISH 1966-08-25
157 Zanthoxylum kauaense Waianae Mts., Kunia trail, just E of Puu Kanehoa ridge Preserved_Specimen 35' tree; fruit green, small Stone, B.C. 3296 Oahu BISH 1960-03-26
158 Zanthoxylum kauaense near Kolekole in remnant patch of intact forest. with Alectryon, Pleomele forbesii, Phyllostegia hirsuta Preserved_Specimen ~6 m pole tree Takeuchi, W.N. 2814 Oahu BISH 1986-09-06
159 Zanthoxylum kauaense Half way up Kaala from Fiore-Break Trail Forest Preserved_Specimen Seedling! Degener, O. 10402 Oahu BISH 1936-02-02
160 Zanthoxylum kauaense Anahulu Trail, Koolau Range rainy Preserved_Specimen Degener, O. 10084 Oahu BISH 1935-06-16
161 Zanthoxylum kauaense Ridge S. of S.Opaeula Gulch, Paalaa, Koolau Mts Open moist woods Preserved_Specimen 15 ft. x 3 in. tree St.John, H. 13339 Oahu BISH 1933-09-24
162 Zanthoxylum kauaense Waianae Mts., trail to Puu Kanehoa N facing slope, open forest Preserved_Specimen 6 m x 13 cm tree; sparsely foliated, mostly at tips of branches; leaflets all 3 Marr, K.L. 37 Oahu BISH 1988-06-07
163 Zanthoxylum kauaense near Kolekole on sliding rubble Preserved_Specimen Takeuchi, W.N. 2815 Oahu BISH 1986-09-06
164 Zanthoxylum kauaense Koaie, Waimea gulch bottom Preserved_Specimen 10 m tree; fruit elliptical Montgomery, S.L. Kauai BISH 1987-06-11
165 Zanthoxylum kauaense Waianae Mts., Palikea near top of gulley below summit; with Pittosporum, Metrosideros, Dubautia, Lobelia yuccoides Preserved_Specimen ~2 m tree; fruiting; leaves with citrus odor Marr, K.L. 65 Oahu BISH 1988-06-26
166 Zanthoxylum kauaense NW boundary of Kaluaa with Zanthoxylum skottsbergii, Alectryon mahoe, Cyanea membranacea, Nothocestrum longifolium, Viola tracheliifolia Preserved_Specimen 4 m x 5 in. Takeuchi, W.N. 2821 Oahu BISH 1978-09-07
167 Zanthoxylum kauaense Near summit of Pupukea-Kahuku trail Exposed rain-forest on ridge Preserved_Specimen Degener, O. 4109 Oahu BISH 1931-12-06
168 Zanthoxylum kauaense Lihue Distric; Top of Mt. Haupu; Metrosideros Lowland Mixed Mesic Forest with North facing Native cliffs; Associated Plants: Bidens valida, Artemisia, Sida fallax, Hedyotis acuminata & H. fluviatilis, Bonamia, Isodendrion, Zanthoxylum, Carex wahuensis, Eragrotis variabilis, Hibiscus kokio, Lobelia niihauensis, Machaerina, Pipturus, Lepidium bidentatum, Mariscus, Melicope anisata. Preserved_Specimen 2.5 m tall tree; in fruit; Wood, K.R. 2037 Kauai BISH 1992-07-27
169 Zanthoxylum kauaense Waianae Range, Palehua In decadent, pastured forest Preserved_Specimen Three trees. Degener, O. 10318 Oahu BISH 1935-11-23
170 Zanthoxylum kauaense land section of Makakupaia, Kupaia Gulch Associates: Antidesma, Pisonia, Ilex, Athyrium, Metrosideros, Coprosma Preserved_Specimen A 4m tall tree growing adjacent to the stream bed of the gulch. The tree appears unhealthy and on the verge of succumbing. Only about 20 percent of the leaves on the tree are compound. Lau, J. 1451 Molokai BISH 1985-09-24
171 Zanthoxylum kauaense Waianae Mts., Puu Kaua N facing ridge of N ridge of gulch of main ridge E of summit.; with Metrosideros, koa, Alyxia, Pelea, Schinus, Lantana, Psychotria, Chamaesyce Preserved_Specimen leaflets 3-11; fruit Marr, K.L. 51 Oahu BISH 1988-06-24
172 Zanthoxylum kauaense Olaa Forest Reseve, behind Volcano dump Wet forest habitat. Preserved_Specimen 13m x 19.5cm. Not a healthy looking tree. Leaflets 3-5, shape reminiscent of trees from Wai'anae Mtns. of Oahu. One other mature nearby (12m x 12cm). Both trees sterile. Marr, K.L. 80 Hawaii BISH 1988-07-30
173 Zanthoxylum kauaense Puu Kaua, Waianae Mts woods near summit Preserved_Specimen Gillespie, J.W. 1064 Oahu BISH 1927-07-06
174 Zanthoxylum kauaense Paalaa South Opaeula Gulch, Koolau Mts on wooded ridge Preserved_Specimen Hosaka, E.Y. 341 Oahu BISH 1930-11-09
175 Zanthoxylum kauaense Kauaikinana Stream, Kokee Above stream on wooded slope Preserved_Specimen Stone, B.C. 3389 Kauai BISH 1960-04-14
176 Zanthoxylum kauaense Mokuleia, makua divide, Waianae Mts rain forest Preserved_Specimen 20' x 3 tree; leaflets thick membranous, not puntate St.John, H. 12235 Oahu BISH 1932-11-25
177 Zanthoxylum kauaense SE of Schofield, Puu Hapapa ridge, forest Preserved_Specimen Degener, O. 17737 Oahu BISH 1939-06-20
178 Zanthoxylum kauaense Waimea District. Kula, upper southern headwater gulch. Metrosideros-Acacia montane mesic forest with Dodonaea viscosa, Psychotria mariniana, P. greenwelliae, Wikstroemia furcata, Kadua affinis, Coprosma waimeae, Antidesma platyphylla hillebrandii, Claoxylon sandwicensis, Melicope barbigera, M. ped Preserved_Specimen Tree, 4 metrs tall, in early bud, several trees in immediate area Wood, K.R. 16604 Kauai BISH 2015-09-13
179 Zanthoxylum kauaense Kawela AHU. Tr 6(1); On ridge betwn W & E forks of Kawela Gulch Mesic 'ohi'a forest with other native trees and a native shrub, matted-fern understory. Preserved_Specimen Small tree, shrubby form; 3m tall Warshauer, F.R. 2312 Molokai BISH 1979-06-20
180 Zanthoxylum kauaense Waianae Mtns. Kaluaa Gulch. Same site as KM 2, 3, 69-71, 73 Other species: Metrosideros, Streblus, Morinda trimera, Antidesma, Acacia koa, Syzygium sandwicensis, Nothocestrum, Alectryon macrococcum, Charpentiera obovata, Pisonia spp, Pipturus albidus, Urera glabra, Pelea spp, Platydesma cornuta. Preserved_Specimen Total of 17 trees this site, 6 in fruit. Three seedlings. 11m x 9cm DBH Marr, K.L. 68 Oahu BISH 1988-07-17
181 Zanthoxylum kauaense Kumueli Gulch On steep slope near stream bed. Vegetation: Metrosideros, Cheirodendron, Perrottetia, Kadua, Urera, Pipturus, Cyrtandra, Sadleria, Psychotria, Cibotium, Pneumatopteris, Diplazium, Athyrium, Ilex, Coprosma, Peperomia, Selaginella, Broussaisia, Psilotum, Tetraplasandra, Pittosporum, Elaphoglossum, Dicranopteris. Preserved_Specimen Few small trees to 4m tall, with saplings nearby. Leaves with a lemon/soapy smell when crushed. Oppenheimer, H.L. 20722 Molokai BISH 2007-02-22
182 Zanthoxylum kauaense Kaluaa, near head of basin forest, with Syzygium, Myrsine, Elaeocarpus Preserved_Specimen 3-4 m tree; all inds. In colony are fertile, good fruiting material present on adjacent trees but unreachable. Takeuchi, W.N. 2818 Oahu BISH 1986-09-07
183 Zanthoxylum kauaense Hapapa canopy break Preserved_Specimen small tree Takeuchi, W.N. 3419 Oahu BISH 1987-02-01
184 Zanthoxylum kauaense North side of Haupu talus slope Preserved_Specimen Small tree. Ht. 2 m. MacDaniels, L.H. 731 Kauai BISH 1927-02-16
185 Zanthoxylum kauaense Kawaiiki Valley, off Kaluahaulu ridge, N facing slope, below the ridge and below the Diellia erecta f. alexandri population Mixed Mesic Forest canopy of Acacia koa and Metrosideros polymorpha w/ Tetraplasandra, Cheirodendron trigynum, Dianella sandwicensis, Poa sandwicensis, Luzula, Rumex, Bidens cosmoides, Dubautia laevigata, Kadua affinis, Cryptocarya mannii, Peperomia. Preserved_Specimen Tree 6 ft tall w/ dead top about 10 ft before steep cliff. Immature green frt. Purple petioles, dark green lvs. About 5 seedlings. Tangalin, N. 1969 Kauai BISH 2009-04-08
186 Zanthoxylum kauaense N head of Kahanahaiki Valley dense forest Preserved_Specimen Degener, O. 20639 Oahu BISH 1950-05-07
187 Zanthoxylum kauaense Waimea District, Kawaiiki, off Kaluakaulu ridge, upper forest and drainage to the south of Koaie and north of Waialae Metrosideros polymorpha mixed mesic forest w/ Dubautia laevigata, Hedyotis terminalis, Cheirodendron trigynum, Dianella sandwicensis, Poa sandwicensis, Bidens cosmoides, Peperomia membranacea Preserved_Specimen Tree, 4 m tall, moderately branched, leaves dark green, with fine puberulence on young stems and leaves, in flower, occasional with Z. dipetalum Wood, K.R. 9019 Kauai BISH 2001-06-27
188 Zanthoxylum kauaense Kamoku, Kaiholena Gulch, SW ridge low forest Preserved_Specimen 4 m x 6 cm tree; wood hard; leaves thick, subcoriaceas St.John, H. 22593 Lanai BISH 1947-04-04
189 Zanthoxylum kauaense ridge crest between Kaua and Kanehoa. field no. Waianae 517a small individual in a Chamaesyce dominated clearing Preserved_Specimen Takeuchi, W.N. 2847 Oahu BISH 1986-09-20
190 Zanthoxylum kauaense Waianae Mts., trail on E ridge of Puu Kaua N facing slope Preserved_Specimen 4 m x 10 cm; leaflets 3-5; sterile Marr, K.L. 22 Oahu BISH 1988-04-17
191 Zanthoxylum kauaense Hapapa, subcrest of main divide forest; with Syzygium, Pelea Preserved_Specimen 8-9' tree Takeuchi, W.N. 3905 Oahu BISH 1988-07-17
192 Zanthoxylum kauaense Waianae Mts., Kaluaa Gulch E facing slope of small gulley in N part of gulch Preserved_Specimen 11 m x 9 cm Marr, K.L. 69 Oahu BISH 1988-07-17
193 Zanthoxylum kauaense S slope of Haleakala, on ridge between E&W forks of Pahihi Gulch Dry exotic grassland with native and exotic shrubs and scattered native trees Preserved_Specimen Warshauer, F.R. 3091 Maui BISH 1980-08-20
194 Zanthoxylum kauaense Waimea Puu Ka Pele FR on W side of Waimea Canyon steep slope, grassy, dry; with Acacia koa, Metrosideros, Myrsine Preserved_Specimen 2.5 cm x 5 m reclining treelet; branches straggling. Corolla lobes pale green, apex and margins purple, anthers yellow, filaments pale green Lorence, D.H. 5234 Kauai BISH 1987-05-15
195 Zanthoxylum kauaense Kipuka Puaulu growing near beginning of trail Preserved_Specimen tree: 20' high appears to be dying Fruit: texture of lime,citmus odor Fagerlund, G.O. 779 Hawaii BISH 1943-08-16
196 Zanthoxylum kauaense Waianae Kai. on saddle ridge near poles with koa, ohia, Grevillea, Dodonaea, maile, pilo, Scaevola Preserved_Specimen 12' tree, with immature seed Perlman, S. 5047 Oahu BISH 1987-01-08
197 Zanthoxylum kauaense Kamaileunu ridge, edge of stepp slopes a local concentration near the electrical transmission towers Preserved_Specimen Takeuchi, W.N. 2129 Oahu BISH 1985-03-31
198 Zanthoxylum kauaense Kawaiiki, off Kaluahaulu ridge, upper forest. Metrosideros-Acacia mixed mesic forest with Dubautia laevigata, Kadua affinis, Cheirodendron spp., Dianella sandwicensis, Poa sandwicensis, Bidens cosmoides, Peperomia membranacea, Microlepia strigosa, Dryopteris fuscoatra, Cyanea leptostegi Preserved_Specimen Tree, 7 meters tall, very large leaves, vigorus, in fruit Wood, K.R. 16544 Kauai BISH 2015-07-14
199 Zanthoxylum kauaense Waianae Mountains. Puu Kaua, beside trail from crest right between Puu Kaua and Pohakea Pass, third minor ridge from summit of Puu Kaua Below small stand of Sophora. Close to individual Zanthoxylum dipetalum with strong anixe odor. Preserved_Specimen Sterile. 4 meters tall x 25 cm DBH. Marr, K.L. 55 Oahu BISH 1988-06-24
200 Zanthoxylum kauaense Kauai Hanalei District. Kahuamaa Flat. Metrosideros-Cheirodendron-Dicranopteris montane wet forest with Kadua affinis, Dodonaea viscosa, Planchonella sandwicensis, Polyscias waimeae, P. waialealae, Melicope clustifolia, Zanthoxylum dipetalum, Ilex anomala,Bobea brevipes, Nestegis sandwicensis, Scaev___ Preserved_Specimen Tree, 4.5 meters tall, stems dark brown, vegetative, mapped. Wood, K.R. 18238 Kauai BISH 2019-07-09
201 Zanthoxylum kauaense not far from summit of Kunia Trail, which ends on summit ridge of Waianae Mts. between Puu Kanehoa and Puu Hapapa On steep slopes in scrubby Metrosideros polymorpha forest, with Styphelia tameiameiae, Antidesma, Diospyros, Alyxia oliviformis, Pelea spp. Preserved_Specimen Rounded shrub 5 feet tall Carlquist, S. 1903 Oahu BISH 1966-07-02
202 Zanthoxylum kauaense Waimea Canyon; mile 11.3 on Hwy. 550 forest, steep dry slope; with Zanthoxylum dipetalum, Myrsine, Acacia, Pelea Preserved_Specimen 15' x 3 tree Flynn, T.W. 2214 Kauai BISH 1987-05-21
203 Zanthoxylum kauaense Waianae Mts., Kaluaa, Glch. Betwn Puu Kanehoa & Puu Hapapa. Central Kaluaa. One gulch N. of main glch. most commonly visited. At fork, nr. gulch head with Tetraplasandra, Claoxylon, Alectryon, Cyanea, Viola, Streblus, Neraudia Preserved_Specimen 3 plants, w/fr Perlman, S. 4926 Oahu BISH 1986-08-07
204 Zanthoxylum kauaense Along highest ridge overlooking Kahuku along Pupukea-Kahuku Trail In rainforest Preserved_Specimen Degener, O. 4109 Oahu BISH 1931-12-06
205 Zanthoxylum kauaense West Maui, Hanaula, Manawainui steep slope near gulch bottom Preserved_Specimen 3-4 m tree; leaves glandular on margin; fruit glandular, 11-12 mm long Hobdy, R.W. 1720 Maui BISH 1982-12-15
206 Zanthoxylum kauaense Pahole Gulch, Waialua, Waianae Mts near crest of steep spur, south seciton of gulch. Canopy of Grevillea and Metrosideros Preserved_Specimen Small tree 20 ft. ht., 3 in. dbh. Corolla 4. Leaflets 5, rarely 3. Flowers pungent. Nagata, K.M. 1541 Oahu BISH 1978-01-31
207 Zanthoxylum kauaense Waimea District. Kawaiiki region, north facing slopes below dividing ridge above twin falls. Metrosideros-Acacia mesic forest with Myrsine lanaiensis, Myoporum sandwicensis, Kadua affinis, Xylosma hawaiiense, Planchonella sandwicensis, Ilex anomala, Perrottetia sandwicensis, Charpentiera elliptica, Chrysodracon aurea, Elaeocar Preserved_Specimen Tree, 4 meters tall, trunk mottled gray, some fruit, mapped. Wood, K.R. 17169 Kauai BISH 2016-11-01
208 Zanthoxylum kauaense East Maui, Makawao District, Makawao FR, Kahakapao drainage basin Growing on windward facing slope in mesic forest. Associated vegetation: Acacia, Metrosideros, Dicranopteris, Psychotria, Melicope, Pleomele, Nestegis, Pouteria, Clermontia, Fraxinus, Rubus. Preserved_Specimen Single 8m tall tree, lvs slightly lemon scented. Oppenheimer, H.L. 70606 Maui BISH 2006-07-12
209 Zanthoxylum kauaense Nuololo, north facing slopes above drainage Metrosideros-Acacia montane mesic forest, 40-50% open, up to 20 m tall canopy, W/ Alphitonia, Pouteria, Xylosma hawaiiensis, Claoxylon, Myrsine lanaiensis, Wikstroemia furcata, Dodonaea, Kadua affinis, Melicope ovata, Pleomele, Tetrplasandra kavaiensis, Psychotria greenwelliae Preserved_Specimen tree, 2-3 m tall, tan-brown bark, 5 cm diameter at base, single tree Wood, K.R. 14541 Kauai BISH 2011-02-28
210 Zanthoxylum kauaense Kaaikukae Gulch, Waianae Mts W facing slope, moist, shaded Preserved_Specimen tree; flower white inside, petals purple Gagné, W.C. 620 Oahu BISH 1971-01-30
211 Zanthoxylum kauaense Waianae Mts., Palikea trail, gulley below summit with Cryptomeria, Schinus Preserved_Specimen 8 m x 16 cm tree; 5 leaflets; leaves with citrus odor; sterile Marr, K.L. 60 Oahu BISH 1988-06-26
212 Zanthoxylum kauaense Waianae Mts., Puu Kaua N facing slope of N ridge of gulch E of summit; with koa, Metrosideros, Alyxia, Schinus, Pelea Preserved_Specimen variation in leaflets 3-11 Marr, K.L. 50 Oahu BISH 1988-06-24
213 Zanthoxylum kauaense Waimea District. Kawaiiki Valley, above Twin Falls area. Diverse Mesic Forest W/Metrosideros-Acacia koa canopy, with Zanthoxylum, Myrsine mezii, Coprosma, Dodonea, Dubautia, Kadua affinis, Syzgium, Dianella, Perrottetia sandwicensis, Claoxylon sandwicensis, Psychortria mari Preserved_Specimen 5.5 ft. tall small tree. Leaflets dull dark green above and dull med-green below w/ dark green vegetation. Fruit med green w/ darker green oil dots. Bark rough grey. Three trees in this area. Tangalin, N. 1198 Kauai BISH 2012-04-13
214 Zanthoxylum kauaense Koolau Range, Malaekahana-Waimea, Summit trail summit woods Preserved_Specimen tree 5 m. x 8 cm. male fl. white. St.John, H. 18666 Oahu BISH 1938-01-09
215 Zanthoxylum kauaense Huliwai mixed native exotic forest, steep dirt slopes; with Alyxia Preserved_Specimen few fruit Takeuchi, W.N. 2845 Oahu BISH 1986-09-20
216 Zanthoxylum kauaense buttress ridge NW of Kaluaa gulch, near summit divide; interval between Puu Hapapa and Kanehoa a local concentration on steep slopes Preserved_Specimen Takeuchi, W.N. 2048 Oahu BISH 1985-03-03
217 Zanthoxylum kauaense Hilo woods Preserved_Specimen Hawaii BISH
218 Zanthoxylum kauaense Mawini makai of forest reserve fence Preserved_Specimen Degener, O. 19310 Maui BISH 1948-12-26
219 Zanthoxylum kauaense Waianae Mts., Puu Kaua, trail on main E ridge from summit N facing slope of S ridge of gulley on E end of ridge Preserved_Specimen sterile; leaflets 3-5 Marr, K.L. 21 Oahu BISH 1988-04-17
220 Zanthoxylum kauaense Waimea District. Makaha valley, 1.3 miles down Makaha Ridge road from Hwy 550, below Kokio Keokeo picnic area Mixed mesophytic forest dominated by Acacia koa with Metrosideros,, Pelea barbigera, Remya kauaiensis and Alphitonia. Preserved_Specimen Tree of 20 ft. Lvs. dark green, dull, with yellow-green midrib, below paler with yellow-green midrib. Some reddish tinting at base of petiolules. Frt. green with darker green glands. Flynn, T.W. 2774 Kauai BISH 1988-02-24
221 Zanthoxylum kauaense Olaa Forest Reserve, north of Volcano dump Rainforest with Cibotium, Koa, Metrosideros, Psychotria. Preserved_Specimen Leaflets mostrly 3, leaves and bark with strong citrus odor, sweet. Some old fruit on tree. Marr, K.L. 33 Hawaii BISH 1988-06-01
222 Zanthoxylum kauaense Koolau Mts., Pupukea-Kahuku trail ridge Preserved_Specimen 25' tree; leaflets 3; fruit green; seeds shiny black Stone, B.C. 2816 Oahu BISH 1959-05-04
223 Zanthoxylum kauaense Iao valley, Nakalaloa gulch Aleurites, Metrosideros, Dicranopteris linearis, Hedyotis acuminata, Psidium cattleianum, Rubus argutus, Freycinetia, Pipturus, Psychotria mariniana, Clermontia kakeana, Xylosma hawaiiense, Alyxia, Charpentiera ovata, Hedyotis terminalis Preserved_Specimen 5 m; fruit green turning purple Wood, K.R. 8987 Maui BISH 2001-05-30
224 Zanthoxylum kauaense Waianae Mts., Puu Hapapa, N of main ridge small valley Preserved_Specimen Storey, W.B. Oahu BISH
225 Zanthoxylum kauaense Olaa Forest Reserve, north of Volcano dump Wet forest habitat. Other species: Ilex, Vaccinium, Rubus (introduced), Tetraplasandra, Cyrtandra, Gouldia, Metrosideros, Pelea, Cheirodendron. Preserved_Specimen 11m x 20cm. Fruit on ground. New flush of leaves and infloresences. Marr, K.L. 78 Hawaii BISH 1988-07-30
226 Zanthoxylum kauaense Along summit trail on right side about 1 mile beyond shelter house at junction of Pupukea & Summit Trail Growing Ilex anomela, Broussaisia spp., Pelea, Suttonia sandwicensis, Gouldia spp., etc. Preserved_Specimen One leaf with 5 leaflets. Pearsall, G.A. 398 Oahu BISH 1959-06-13
227 Zanthoxylum kauaense Halemanu, above Waimea Rather dry situation in side of valley Preserved_Specimen Tree 4 m. high MacDaniels, L.H. 767 Kauai BISH 1927-02-16
228 Zanthoxylum kauaense middle Palawai Ridge Dark forest Preserved_Specimen Degener, O. 10864 Oahu BISH 1936-05-12
229 Zanthoxylum kauaense Lanai trails Preserved_Specimen transitory form Munro, G.C. 236 Lanai BISH 1915-06-19
230 Zanthoxylum kauaense Malaekahana Trail, Laie (Near No. 10083) Forested ridge half way to summit Preserved_Specimen Degener, O. 10086 Oahu BISH 1935-07-29
231 Zanthoxylum kauaense Kalalau Lookout forest Preserved_Specimen Degener, O. 20599 Kauai BISH 1950-02-07
232 Zanthoxylum kauaense Puu Kanehoa, Waianae Range dry wooded ridge Preserved_Specimen tree Hume, E.P. 487 Oahu BISH 1932-01-18
233 Zanthoxylum kauaense Olaa Forest Reseve, north of Volcano dump Rainforest with Cibotium, Koa, Metrosideros, Psychotria. Preserved_Specimen 7m tall x 16cm DBH. Leaflets 3-5, infected possibly with insect. No infloresence. Leaves and bark with sweet, citrus odor. Marr, K.L. 34 Hawaii BISH 1988-06-01
234 Zanthoxylum kauaense Kokee, Halemanu forest Preserved_Specimen Degener, O. 28097 Kauai BISH 1960-11-15
235 Zanthoxylum kauaense Puu Hapapa, S ridge buttress crumbling rock; with Alectryon, Streblus, Pisonia Preserved_Specimen 4 m Takeuchi, W.N. 2800 Oahu BISH 1986-09-01
236 Zanthoxylum kauaense Kaluaa, near head of basin forest, with Syzygium, Myrsine, Elaeocarpus Preserved_Specimen 3-4 m tree; all inds. In colony are fertile, good fruiting material present on adjacent trees but unreachable. Takeuchi, W.N. 2818 Oahu BISH 1986-09-07
237 Zanthoxylum kauaense Waianae Mts., Palikea Trail protected hillside Preserved_Specimen ~5' bush; flowers fragrant; calyx green, edged with white; corollar white; ovary and style green, stigma white; filaments white, anthers brown Crosby, M.R. 1918 Oahu BISH 1964-07-31
238 Zanthoxylum kauaense Skyline Drive, Ulupalakua dry, decadent forest Preserved_Specimen Degener, O. 28095 Maui BISH 1960-01-17
239 Zanthoxylum kauaense Waianae Mts, Puu Kalena. On S. side of Puu Kalena in gully off E. side of Ridge Tr to summit With other trees Nothocestrum Iongifolium, Charpentiera tomentosa, Labordia tinifolia, Psychotria Preserved_Specimen 8 plants seen. Flowers white Perlman, S. 4900 Oahu BISH 1986-07-27
240 Zanthoxylum kauaense W Maui, headwater region of Honokahua Valley. Riparian zone of ohia montane mesic forest within Unit 4 of Puu Kukui Watershed Mgmnt Area On E bank 15 ft. above intermittent stream. Assoc. with Dicranopteris linearis, Metrosideros polymorpha, Pipturus albidus, Tibouchina herbacea, Cheirodendron trigynum, Bobea elatior, Psychotria mariniana, Clermontia arborescens, Ageratina adenophorus, Nothocestrum longifolium, Cyanea scabra, C. macrostegia, Coniogramme pilosa, Amauropelta globulifera, Peperomia Preserved_Specimen Ca. 95% foliated; appears to be senescent & in poor health. Single specimen found. Meidell, J.S. 100 Maui BISH 1995-10-18
241 Zanthoxylum kauaense Kopiwai trail, Kokee rain forest Preserved_Specimen 18' foot tree Degener, O. 21569 Kauai BISH 1952-01-11
242 Zanthoxylum kauaense Piko Trail, Makua Valley, Kawaihapai forest at top Preserved_Specimen tree Degener, O. 8597 Oahu BISH 1932-07-10
243 Zanthoxylum kauaense Waimea canyon, Kawaliki drainage area In Acacia koa- Metrosideros mesic forest with Styphelia, Dianella sandwicensis, Psychotria mariniana, Santalum freycinetianum var pyrularium, Charpentiera elliptica, Claoxylon, Lantana, Rubus argutus, Psidium cattleianum. Preserved_Specimen Small tree 15 ft ht with immature frt, leaflets thick, coriaceous, ovate, glabrous, margins entire, apex acute. Perlman, S. 19792 Kauai BISH 2005-09-23
244 Zanthoxylum kauaense Waianae range, W of Camp Kunia with Pelea, Gouldia, Pittosporum Preserved_Specimen 10' spreading tree Wuotila, J.L. Oahu BISH 1958-12-18
245 Zanthoxylum kauaense Ekahanui-Kaua among boulder rocks; in Pelea Preserved_Specimen 4 m tree Takeuchi, W.N. 3554 Oahu BISH 1987-05-16
246 Zanthoxylum kauaense North Kona District: Puuwaawaa, Waiho. 1/2 mile NNW of Hibiscadelphus grove Hidden in mamane trees. Preserved_Specimen One tree. 6 inches in diameter, 20 feet high. Male flower in parts of 4. Corolla white inside, purplish outside. Leaves: light green, dull; veins red above. Fagerlund, G.O. 1016 Hawaii BISH 1945-03-24
247 Zanthoxylum kauaense Waimea District. Makaha valley, 1.3 miles down Makaha Ridge road from Hwy 550, below Kokio Keokeo picnic area Mixed mesophytic forest dominated by Acacia koa with Metrosideros,, Pelea barbigera, Remya kauaiensis and Alphitonia. Preserved_Specimen Tree of 20 ft. Lvs. dark green, dull, with yellow-green midrib, below paler with yellow-green midrib. Some reddish tinting at base of petiolules. Frt. green with darker green glands. Flynn, T.W. 2774 Kauai BISH 1988-02-24
248 Zanthoxylum kauaense Waimea District. Makaha, south facing slopes below Milolii road. Acacia-Metrosideros lowland mesic forest and steep slopes of Dicranopteris with Alphitonia ponderosa, Kadua affinis, Planchonella sandwicensis, Caloxylon sandwicensis, Coprosma waimeae, Polyscias kavaiensis, Melicope ovata, Melicope barbigera, Dodonaea viscosa, Nestegis sandwicensis. Preserved_Specimen Tree, 4 meters tall, trunk mottled gray-brown wit immature fruit, 2 trees in immediate area, mapped. Wood, K.R. 17084 Kauai BISH 2016-07-28
249 Zanthoxylum kauaense Honouliuli contour mauka slopes; mesic forest Preserved_Specimen shade leaves detached at node; usually trifolate, but occasional 5-, 7- pinnate Takeuchi, W.N. 3579 Oahu BISH 1987-05-31
250 Zanthoxylum kauaense Puna DISTR. 'Olaa AHU. Tr 32(95/96); E slope of Mauna Loa near Volcano dump; N of Hwy 11 Wet 'ohi'a forest with other native trees and a treefern and native shrub understory. Preserved_Specimen 2cm Bd; 10m tall. Unhealthy appearance. Warshauer, F.R. 1256 Hawaii BISH 1977-07-23
251 Zanthoxylum kauaense Mt. Kaala forest Preserved_Specimen 13' tree Degener, O. 19719 Oahu BISH 1949-07-10
252 Zanthoxylum kauaense W Maui, S facing wall of N fork of Poahakea located below hill marked 3896 on USGS Wailuku quad. Gulch, accessed from dead-end spur to the south of pasture Consistent with `ohi`a lowland mesic forest. Assoc. veg. Metrosideros polymorpha, Perrottetia sandwicensis, Coprosma foliosa, Psychotria mariniana, Ilex anomala, Hedyotis terminalis, Smilax melastomifolia, Lythrum maritimum, Cuphea carthagenensis, Dicranopteris linearis, Microlepia strigosa, Sadleria cyatheoides. Preserved_Specimen In good health, fully foliated & ca. 14 ft high; gray-brown bark conspicuously glandualar punctate. Foliage w/strong lemon scent. Meidell, J.S. 104 Maui BISH 1996-01-22
253 Zanthoxylum kauaense W Maui, Lahaina District, headwall of Ukumehame Valley, SW of the Hanaula Reservoirs On steep slope. Preserved_Specimen Uncommon, tree of 4 m, healthy. Oppenheimer, H.L. 59802 Maui BISH 1999-05-14
254 Zanthoxylum kauaense Waimea district. Kawaiiki, northern slopes below Kaluahaulu ridge. Metrosideros-Acacia mixed mesic forest w/ Dodonaea, Santalum freycinetianum, Kadua affinis, Xylosma hawaiiensis, Pouteria sandwicensis, Antidesmia platyphylla, dubautia laevigata, wikstroemis oahuensis, Zanthoxylum dipetalum, Melicope ovata Preserved_Specimen Tree, 3 m tall, trunk gray, with fruit, capsules dark green, seed gloosy black, mapped. Wood, K.R. 16309 Kauai BISH 2015-02-11
255 Zanthoxylum kauaense Kapua Gulch, S branch forest Preserved_Specimen Degener, O. 28530 Kauai BISH 1963-08-25
256 Zanthoxylum kauaense Kokee stream woods Preserved_Specimen 12' leafy tree Degener, O. 23894 Kauai BISH 1955-09-14
257 Zanthoxylum kauaense Hono O Na Pali NAR, Pohakea region, steep ridge above Hanakapiai Open Metrosideros mesic forest with adjacent transitional wet forest. W/ Kadua affinis, Dicranopteris, Myrsine linearifolia, Scaevola procera, Xylosma hawaiiense, Cyrtandra paludosa, Lysimachia sp, Santalum pyrularium, Dubautia laevigata, Psychotria mariniana, Coprosma Preserved_Specimen Tree, 3.5 m tall, 2-trunked, mottled gray-brown with light green lichen, in flower, single tree Wood, K.R. 14728 Kauai BISH 2011-07-28
258 Zanthoxylum kauaense West Maui, Lahaina, Kauaula Valley Moderately steep, windward facing upper to mid slope; mesic/dry forest and shrubland mosaic. Associated vegetation: Dodonaea, Dubautia linearis, Diospyros, Metrosideros, Remya, Psychotria mauiensis, Achyranthes splendens (100's of plants on this slope), Lipochaeta connata, Psydrax, Senna, Pleomele, Myrsine, Bidens micrantha, Sophora, Alyxia, Dryopteris sandwicensis, Nestegis, Asplenium adiantum-nigrum, Coprosma foliosa, Osteomeles, Wikstroemia, Leptecophylla. Preserved_Specimen 3 m tree. Oppenheimer, H.L. 10605 Maui BISH 2006-01-28
259 Zanthoxylum kauaense ridge flute near Kamakalii native forest; with Pelea oahuensis Preserved_Specimen Takeuchi, W.N. 2826 Oahu BISH 1986-09-01
260 Zanthoxylum kauaense near head of Kaluaa base mesic forest Preserved_Specimen 4-5 m tree Takeuchi, W.N. 3710 Oahu BISH 1987-11-08
261 Zanthoxylum kauaense Huliwai a very large leaved tree growing next to Zanthoxylum Preserved_Specimen Takeuchi, W.N. 2846 Oahu BISH 1986-09-20
262 Zanthoxylum kauaense Paalaa South Opaeula Gulch, Koolau Mts on wooded ridge Preserved_Specimen Hosaka, E.Y. 341 Oahu BISH 1930-11-09
263 Zanthoxylum kauaense Kopakapaka ridge, S of Makaha, north fork of Kauhau Acacia-Metrosideros mesic forest. W/ Alphitonia, Dodonaea, Santalum freycinetianum, Antidesma, Nestegis, Leptecophylla, Melicope ovata, M. barbigera, Pouteria, terrestrial cover of Carex wahuensis, C. meyenii, Gahnia beecheyi, Dianell Preserved_Specimen tree, 6 m tall, trunk gray-brown mottled, vigorous, with fruit, uncommon, ca. 10 plants observed during day in region Wood, K.R. 13852 Kauai BISH 2009-07-23
264 Zanthoxylum kauaense Waimea Canyon, toward head from Puu Kapele dryland forest Preserved_Specimen 12' x 5 tree; inodorous Christensen, C. 199 Kauai BISH 1977-05-07
265 Zanthoxylum kauaense Waianae Mts., Palikea trail, ~500 m N of peak S facing slope Preserved_Specimen 7 m x 3 cm tree; many fruit; leaves with citrus odor Marr, K.L. 62 Oahu BISH 1988-06-26
266 Zanthoxylum kauaense Waianae Mtns. Puu Kaua north ridge of gulley off of large main ridge directly east of summit on north facing slope At this site 10 other indiviuals, at least 3 with fruit (KM 50 & 51). Preserved_Specimen Marr, K.L. 75 Oahu BISH 1988-06-17
267 Zanthoxylum kauaense Mt. Kaala, Waianaeuka Metrosideros Preserved_Specimen 4 m x 5 cm shrub St.John, H. 23696 Oahu BISH 1949-05-22
268 Zanthoxylum kauaense Kokee stream forest Preserved_Specimen ~9-12' tree Degener, O. 21484 Kauai BISH 1951-12-31
269 Zanthoxylum kauaense Huliwai a very large leaved tree growing next to Zanthoxylum Preserved_Specimen Takeuchi, W.N. 2846 Oahu BISH 1986-09-20
270 Zanthoxylum kauaense West Maui, Iao Valley, along back saddle of Lao Valley on border with Olowalu Valley in small bowl below ridge, Aspect: NE. Wet, windswept shrubland, steep; scattered individuals in back of bowl. Wety shrubland: Dicranopteris, Lysimachia, Smilax, Vaccinium, Machaerina, Sadleria, Nephroia, Metrosideros, Stenogyne, Lobelia, Euphorb ia, Luzula, Myrsine, Ilex, Cheirodendron Preserved_Specimen small trees, 1-2 meters tall with few branches. Pezzillo, Z. 1034 Maui BISH 2024-11-25
271 Zanthoxylum kauaense Koolau Range, Summit Trail, Malaekahana-Waimea low forest on summit Preserved_Specimen tree 6 m. x 1 dm. St.John, H. 18665 Oahu BISH 1938-01-09
272 Zanthoxylum kauaense ridge W of Kawaikoa Stream dry lower edge of forest Preserved_Specimen 8 m x 15 cm tree St.John, H. 22700 Kauai BISH 1947-08-10
273 Zanthoxylum kauaense Waimea District. Kohua Ridge and south facing slopes above Koaie. Metrosideros-Acacia montane mesic forest w/ Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bob Preserved_Specimen Tree, 5 m tall, vigorous, mostly vegetative, mapped. Wood, K.R. 17177 Kauai BISH 2016-11-03
274 Zanthoxylum kauaense Kokee. Awaawapuhi Trail Beside trail. Preserved_Specimen Single tree 5 cm. x 5 m. Leaflets 3-5, mostly 5. Sterile this date as well as 14.vi.1988. Marr, K.L. 0.6 Kauai BISH 1988-03-25
275 Zanthoxylum kauaense Waianae Mts., below Puu Kaua, Ekahanui Gulch, N aspect In Acacia koa-Metrosideros mesic forest with Urera kaalae, Delissea subcordata, Schiedea kaalae, Alyxia, Cyanea grimesiana subsp. obatae, Dubautia plantaginea, Rumex albescens, Pouteria sandwicensis, Chamaesyce multiformis, Zanthoxylum kauense, Melinis minutiflora, Ageratina riparia, Argeratina adenophora, Rubus rosifolius. Preserved_Specimen 10 m tall with Zanthoxylum diptalum Wood, K.R. 2923 Oahu BISH 1994-01-11
276 Zanthoxylum kauaense Waianae Mts., N branch of Kaulaa Gulch, S of Puu Hapapa E facing slope at head of side gulch Preserved_Specimen sterile Marr, K.L. 3 Oahu BISH 1988-01-10
277 Zanthoxylum kauaense Kahanahaiki gulch side; N sheltered slope Preserved_Specimen Judd, C.S. 73 Oahu BISH 1928-03-29
278 Zanthoxylum kauaense E slope of Puu Kaua woods Preserved_Specimen 5' tree Degener, O. 28083 Oahu BISH 1962-08-29
279 Zanthoxylum kauaense Kanehoa ridge 4m tree on slopes just below crestline Preserved_Specimen an uncommon sp., but with several conspicuous inds. easily seen from the trail only this specimen fertile Takeuchi, W.N. 1852 Oahu BISH 1984-11-18
280 Zanthoxylum kauaense central ridge of Puu Kanehoa, Waianae Mts., Honouliuli woods Preserved_Specimen 25' x 10 tree St.John, H. 14036 Oahu BISH 1934-01-07
281 Zanthoxylum kauaense Kuaokala with Planchonella, Psidium, Schinus Preserved_Specimen shrub sized Takeuchi, W.N. 2904 Oahu BISH 1986-08-01
282 Zanthoxylum kauaense Waimea District. Kohua Ridge, north facing slopes. Metrosidero-Acacia montane mesic forest w/ Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Kadua affinis, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, M. alyxifolia, Cheirodendron trigynum, Nestegis sandwi Preserved_Specimen Tree, 4 mtall, w/ flower, single tree. Wood, K.R. 18740 Kauai BISH 2021-06-16
283 Zanthoxylum kauaense Puhielelu Ridge, N, valley valley Preserved_Specimen ~8' tree; leaves coriaceous, dull, dark green with yellow midrib and side veins. Inflorescence coming out below leaves Flynn, T.W. 206 Lanai BISH 1982-05-26
284 Zanthoxylum kauaense West Maui, Lahaina District, slopes of Helu, Puehuehunui, first Gulch NW of Luakoi Ridge On S facing slope. Vegetation: Metrosideros, Cheirodendron, Coprosma, Broussaisia, Clermontia, Santalum, Alyxia, Freycinetia, Microlepia, Perrottetia, Diplazium, Ilex, Sadleria. Preserved_Specimen Few trees to 5 m. tall. Oppenheimer, H.L. 50709 Maui BISH 2007-05-07
285 Zanthoxylum kauaense Kokee forest Preserved_Specimen Degener, O. 28098 Kauai BISH 1960-11-19
286 Zanthoxylum kauaense Waianae Mts., Waianae Kai, ridge N of Puu Kawiwi with Rumex, ohia, Hibiscus, koa, Schinus, Dracaena, Rubus Preserved_Specimen 6 m x 15 cm tree; 3-5 leaflets Marr, K.L. 67 Oahu BISH 1988-07-04
287 Zanthoxylum kauaense Awaawapuhi Trail, Honopu Moist Metrosideros woods Preserved_Specimen Tree 10 m. x 8 cm.; perianth white, fl. fragrant St.John, H. 26012 Kauai BISH 1956-12-29
288 Zanthoxylum kauaense Kawaiiki Ridge and nearby drainages to north Metrosideros - Acacia montane mesic forest Alphitonia ponderosa, Santalum pyrularium, Kadua affins, Coprosma waimeae, Ilex anomala, Cheirodendron trigynum, Claoxylon sandwicensis, Scaevola procera, Dodonaea viscosa, Dubautia laevigata, Bidens Preserved_Specimen tree, 3 m tall, trunk gray, in early flower with older fruit, along ridge line, occasional Wood, K.R. 15405 Kauai BISH 2013-02-15
289 Zanthoxylum kauaense Waianae Mts., S gulch between Puu Hapapa and Puu Kanehoa open, N facing slope, 40 m above bottom Preserved_Specimen 6 m x 10 cm; leaflets 3-7; broken leaves have strong odor Marr, K.L. 28 Oahu BISH 1988-05-14
290 Zanthoxylum kauaense Laie-Malaekahana Ridge, Koolau Mts woods Preserved_Specimen 8' young trees St.John, H. 11533 Oahu BISH 1932-02-14
291 Zanthoxylum kauaense Waianae Mts., along ridge extending downward from Puu Hapapa. Entomological collection area code: P-23 Associated with Dracaena and Tetraplasandra. Preserved_Specimen Gagné, W.C. Oahu BISH 1970-04-26
292 Zanthoxylum kauaense upper Awaawapuhi In wet forest dominated by uluhe, ohia, manono, pilo Preserved_Specimen 4 m emergent from uluhe thicket, scattered individuals. Similar to Oahu's skottsbergii but with foliage less variable. Takeuchi, W.N. 3689 Kauai BISH 1987-10-25
293 Zanthoxylum kauaense Koele side of summit ridge, along Munro Trail Metrosideros polymorpha, Dicranopteris mats, Broussaisia arguta, Scaevola chamissoniana, Pipturus rockii, Pittosporum confertiflorum Preserved_Specimen ~5' scraggly shrub Carlquist, S. 2015 Lanai BISH 1966-07-16
294 Zanthoxylum kauaense North Palawai Gulch just north of Palikea, Southern Waianae Range Under a canopy of taller trees, sloping terrain, mesic-summer, dry habitat; three trees, voucher pistillate Preserved_Specimen Obata, J.K. 427 Oahu BISH 1980-03-30
295 Zanthoxylum kauaense Palikea trail, Waianae Mts small valley Preserved_Specimen 3 m; 1 green seed Gagné, W.C. 504 Oahu BISH 1970-04-18
296 Zanthoxylum kauaense N slope of Kaala forest Preserved_Specimen tree Degener, O. 19438 Oahu BISH 1949-02-22
297 Zanthoxylum kauaense ridge near the northermost margin of the Kaluaa drainage occuring in diverse native forest Preserved_Specimen Takeuchi, W.N. 2749 Oahu BISH 1986-08-16
298 Zanthoxylum kauaense Waianae Mts., Palikea trail, SE of summit, below trail with Cryptomeria, Athyrium, Cibotium, Pelea, Perrottetia, Antidesma, Dodonaea, ohia, Coprosma, Psidium, Rubus Preserved_Specimen 4 m x 4 cm tree; leaves with strong odor; sterile Marr, K.L. 56 Oahu BISH 1988-06-26
299 Zanthoxylum kauaense Waianae Mts., Palikea gulley below summit; with Cryptomeria; on eroded bank Preserved_Specimen 3 leaflets, wrinkled; leaves with citrus odor Marr, K.L. 61 Oahu BISH 1988-06-26
300 Zanthoxylum kauaense Waianae Mts., Kaluaa Gulch E facing slope of small gulley in N part of gulch Preserved_Specimen 8 m x 18 cm Marr, K.L. 70 Oahu BISH 1988-07-17
301 Zanthoxylum kauaense Waimea District. Waialae, headwater streamlet just south of Kaluahaulu Ridge Trail. Acacia-Metrosideros montane mesic forest w/ Alphitonia pondersa, Pouteria sandwicensis, Diospyros sandwicensis, Santalum pyrularium, Chrysodracon aurea, Pittosporum glabrum, Cheirodendron trigynum, Ilex anomala, Wikstroemia oahuensis, Claox Preserved_Specimen Tree, 5 m tall, trunk gray-brown with lenticels, in early fruit, 2 trees in immediate area. Wood, K.R. 17521 Kauai BISH 2017-09-05
302 Zanthoxylum kauaense Halfway up Kaala from Fire-Break Trail Forest Preserved_Specimen Degener, O. 10402 Oahu BISH 1936-02-02
303 Zanthoxylum kauaense Waimea Drainage Basin, W side Preserved_Specimen Forbes, C.N. 871 Kauai BISH 1917-07-03
304 Zanthoxylum kauaense East Maui, East Kaupo Gap within dry forest exclusion in Haleakala National Park South slope cliffs. Preserved_Specimen Medeiros, A.C. 214 Maui BISH 1982-07-10
305 Zanthoxylum kauaense Near Kawaihapai steep slopes, dry forest; with Pleomele forbesii, Coprosma fauriei Preserved_Specimen Takeuchi, W.N. Oahu BISH 1984-01-01
306 Zanthoxylum kauaense Waianae Mts., Palikea trail, SE of summit Preserved_Specimen 5 m x 6 cm tree; 3 leaflets; dark green above, lighter below Marr, K.L. 57 Oahu BISH 1988-06-26
307 Zanthoxylum kauaense Hauola Preserved_Specimen Munro, G.C. 441 Lanai BISH 1915-06-09
308 Zanthoxylum kauaense Honouliuli Contour mesic slopes, with Planchonella Preserved_Specimen Takeuchi, W.N. Oahu BISH 1987-02-22
309 Zanthoxylum kauaense Ekahanui Gulch, ascending ridge to Puu Kaua, ridge top, at junction of last adjoining sub-ridge to the ascending ridge associated with Psychotria, Pelea, Koa, etc. Preserved_Specimen Obata, J.K. Oahu BISH 1984-12-23
310 Zanthoxylum kauaense Waianae Mts., Puu Hapapa, N of main ridge small valley Preserved_Specimen Storey, W.B. Oahu BISH 1933-07-30
311 Zanthoxylum kauaense Kaluaa, near head of basin Preserved_Specimen in fruit Takeuchi, W.N. 2904 Oahu BISH 1986-10-11
312 Zanthoxylum kauaense central ridge of Wailupe Valley, Koolau Range rainforest; with Metrosideros polymorpha, Gleichenia linearis Preserved_Specimen Hatheway, W.H. 513 Oahu BISH 1950-12-24
313 Zanthoxylum kauaense Huliwai Preserved_Specimen large leaved tree Takeuchi, W.N. 2846 Oahu BISH 1986-09-20
314 Zanthoxylum kauaense Puu Hapapa, ridge buttress Preserved_Specimen 4 m tree, polelike Takeuchi, W.N. 2804 Oahu BISH 1986-09-01
315 Zanthoxylum kauaense Palikea Trail Preserved_Specimen Bennett, T. 223 Oahu BISH 1981-03-11
316 Zanthoxylum kauaense Aa flow, Devil County road, Keauhou #2 Mauka, U.S.G.S> Quadrangle (Puu Lehua) Preserved_Specimen Tree 12' high, 5' to first limb.Trunk ca. 4 in diameter. Leaves1,2, or 3-foliate. crushed leaves and broken wood with aromatic oil fragrance Gillett, G.W. 1733 Hawaii BISH 1965-04-19
317 Zanthoxylum kauaense Paalaa, Koolau Mts marsh Preserved_Specimen Krauss, N.H. Oahu BISH 1933-12-10
318 Zanthoxylum kauaense Oahu Preserved_Specimen It bore neither fls. nor fruit. Park, K.K. 10159 Oahu BISH 1936-01-05
319 Zanthoxylum kauaense Kalaupapa; Waikolu, S, stream head of valley Preserved_Specimen small to medium tree Hughes, G.D. 1100 Molokai BISH 2004-11-23
320 Zanthoxylum kauaense ridge behind Papaa, Moloaa FR Preserved_Specimen Degener, O. 28096 Kauai BISH 1952-01-16
321 Zanthoxylum kauaense Waianae Mts., Palikea trail, ~500 m N of peak Preserved_Specimen leaves with citrus odor; sterile; trees in fruit Marr, K.L. 63 Oahu BISH 1988-06-26
322 Zanthoxylum kauaense Waianae Mts., Kaluaa Gulch Preserved_Specimen 10 m x 17.5 cm tree; total of 17 trees; 6 in fruit; 3 seedlings Marr, K.L. 73 Oahu BISH 1988-07-17
323 Zanthoxylum kauaense Kaluapuhi Trail, Kokee State Park. Preserved_Specimen Tree, 20' high, 3 dia. (wood from main fork of trunk). Flowers white. Leaves leathery. Fruits still green with dark colored glands. Wood yellowish. Stern, W.L. 3154 Kauai BISH 1974-04-06
324 Zanthoxylum kauaense Waianae Mtns. Kaluaa Gulch. East facing slope on small gulley in north part of the gulch. Same site as KM 2,3, 68-70, 73 Preserved_Specimen Many fruit on this individual, mature. 11m x 37cm DBH Marr, K.L. 71 Oahu BISH 1988-07-17
325 Zanthoxylum kauaense Kanehoa Preserved_Specimen 7-8 m tree with exceptional flower production, scent intense and pervasive and immediately distinguishable from other-genera associates. The largest ind. I have seen; crown massive and spreading; with rounded form. Takeuchi, W.N. 3435 Oahu BISH 1987-02-22
326 Zanthoxylum kauaense Ridge to Kaua Preserved_Specimen Single tree to 4 meters in height; trifoliate leaves Takeuchi, W.N. 3640 Oahu BISH 1987-07-25
327 Zanthoxylum kauaense Waianae Mtns. North branch of Kaluaa Gulch, at head of side gulch, south of Puu Hapapa Preserved_Specimen Stand of 6 trees. Immature fruit. Marr, K.L. 2 Oahu BISH 1988-01-10
328 Zanthoxylum kauaense Puuwaawaa hill Preserved_Specimen Rock, J.F.C. 10206 Hawaii BISH 1912-03-01
329 Zanthoxylum kauaense Auhi, Ulupalakua Preserved_Specimen Munro, G.C. 406 Maui BISH 1915-02-12
330 Zanthoxylum kauaense Waianae Mtns. Palikea. Bottom of gulch leading to summit in Cryptomeria stand Preserved_Specimen All leaves three leaflets, mostly new growth, light green. Trunk split near base. Leaves with citrus odor. Sterile 4m x 16 cm diameter Marr, K.L. 59 Oahu BISH 1988-06-26
331 Zanthoxylum kauaense Pupukea-Kahuku Trail, Kahuku, Koolau Mts. along divide Preserved_Specimen 15 ft. tree, 3 in. in diameter St.John, H. 11099 Oahu BISH 1931-05-24
332 Zanthoxylum kauaense Lanaihale Preserved_Specimen Rock, J.F.C. 8217 Lanai BISH 1910-08-01
333 Zanthoxylum kauaense Kamalo Gulch, above the falls Preserved_Specimen Forbes, C.N. 167 Molokai BISH 1912-07-01
334 Zanthoxylum kauaense Haleakala, Makawao, Waihou Gulch Preserved_Specimen Rock, J.F.C. 10200 Maui BISH 1912-03-01
335 Zanthoxylum kauaense Kokee, jct. of Halemanu and Park Rds Preserved_Specimen small, young tree; bark greyish; foliage dull, coriaceous, not thick, medium grey green above, pale below Herbst, D.R. 1037 Kauai BISH 1968-04-07
336 Zanthoxylum kauaense Rim of Waimea Canyon, below road approx. 11.8 miles from Waimea. Same location as 5 Z. dipetalum individuals Preserved_Specimen Sterile this date. with staminate flowers 13-vi-1988. Most of trunk snapped at base, barely alive this date but looking more vigorous 13-vi-88. Marr, K.L. 0.11 Kauai BISH 1988-03-26
337 Zanthoxylum kauaense Keauhou Mauka: Koa Niu Preserved_Specimen Female tree ion mature fruit. Tree in bad shape; developing lateral branches may keep it alive. Herbst, D.R. 2139 Hawaii BISH 1971-09-16
338 Zanthoxylum kauaense Kanehoa Preserved_Specimen 7-8 m tree with exceptional flower production, scent intense and pervasive and immediately distinguishable from other-genera associates. The largest ind. I have seen; crown massive and spreading; with rounded form. Takeuchi, W.N. 3435 Oahu BISH 1987-02-22
339 Zanthoxylum kauaense Makaha Valley, on steep slope of ridge leading down into Makaha Valley from near well know Pinnacle Rocks, Waianae,Kai area, east of Kamaileunu ridge Preserved_Specimen two trees, 15 ft, height, young fr. Perlman, S. 5058 Oahu BISH 1987-01-08
340 Zanthoxylum kauaense Olaa Forest Preserve, north of Volcanoe dump Preserved_Specimen young infloresence. Leaflets mostly 5. Leaves and stem with citrus odor. 17m tall x 20 cm DBH Marr, M. 31 Hawaii BISH 1988-05-31
341 Zanthoxylum kauaense Kokee, jct. Kalalau Rd. and Kalua-Puhi trail Preserved_Specimen small tree; smooth light bark; dull green foliage and fruit; shiny black seeds Herbst, D.R. 1194 Kauai BISH 1968-08-10
342 Zanthoxylum kauaense Waianae Mts., Kunia trail just E of Puu Kanehoa Preserved_Specimen Stone, B.C. 3273 Oahu BISH 1960-03-26
343 Zanthoxylum kauaense Koele, mountains near Preserved_Specimen Forbes, C.N. 209 Lanai BISH 1913-06-01
344 Zanthoxylum kauaense Ridge to Kaua Preserved_Specimen Single tree to 4 meters in height; trifoliate leaves Takeuchi, W.N. 3640 Oahu BISH 1987-07-25
345 Zanthoxylum kauaense Kaluaa, near head of basin Preserved_Specimen small tree, unisexual and perfect flowers on same ind., all flowers fragrant. Takeuchi, W.N. 2819 Oahu BISH 1986-09-07
346 Zanthoxylum kauaense Papoaea, N slope of Haleakala Preserved_Specimen Forbes, C.N. 2519 Maui BISH 1920-06-14
347 Zanthoxylum kauaense Bird Park Preserved_Specimen Morley, H. 190 Hawaii BISH 1935-04-12
348 Zanthoxylum kauaense Pupukea-Kahuku Trail, Kaumala, Waimea Dry ridge- along trail Preserved_Specimen Ota, M. Oahu BISH 1937-02-19
349 Zanthoxylum kauaense East Maui, Wailou spring area,Olinda on slopes, near gulch bottom Preserved_Specimen A 15' tree growing in shade of ash forest, in bud and flower Hobdy, R.W. 795 Maui BISH 1980-04-20
350 Zanthoxylum kauaense Kaluaa, near head of basin Preserved_Specimen small tree, unisexual and perfect flowers on same ind., all flowers fragrant. Takeuchi, W.N. 2819 Oahu BISH 1986-09-07
351 Zanthoxylum kauaense NE of Puu Hapapa small valley, forest Preserved_Specimen Degener, O. 8599 Oahu BISH 1932-08-07
352 Zanthoxylum kauaense Mauka of Mcgregory Point in open rain forest Preserved_Specimen Degener, O. 22057 Maui BISH 1952-03-09
353 Zanthoxylum kauaense Puuaulu Preserved_Specimen Forbes, C.N. 1031 Hawaii BISH 1915-07-01
354 Zanthoxylum kauaense Koele mountains Preserved_Specimen Forbes, C.N. 113 Lanai BISH 1913-06-01
355 Zanthoxylum kauaense East Maui, Haleakala, E. Kaupo gap steep ridge with dryland forest Preserved_Specimen Crivellone, C. Maui BISH 1983-06-10
356 Zanthoxylum kauaense Hapapa off crest Preserved_Specimen Takeuchi, W.N. Oahu BISH 1987-03-01
357 Zanthoxylum kauaense Puuwaawaa Preserved_Specimen Rock, J.F.C. 3651 Hawaii BISH 1909-06-17
358 Zanthoxylum kauaense slopes of Mt. Kaala Preserved_Specimen 4 m tree Takeuchi, W.N. Oahu BISH 1982-12-01
359 Zanthoxylum kauaense Mahana Preserved_Specimen Munro, G.C. 458 Lanai BISH 1915-07-19
360 Zanthoxylum kauaense Lanaihale Preserved_Specimen Rock, J.F.C. 8217 Lanai BISH 1910-08-01
361 Zanthoxylum kauaense Puuwaawaa Preserved_Specimen Forbes, C.N. 61 Hawaii BISH 1911-06-08
362 Zanthoxylum kauaense Waianae Palehua Preserved_Specimen Skottsberg, C.J.F. 318 Oahu BISH 1922-08-23
363 Zanthoxylum kauaense Honouliuli, Palawai Gulch Preserved_Specimen Tree. Russ, G.W. Oahu BISH 1936-10-19
364 Zanthoxylum kauaense Makaleha Preserved_Specimen Russ, G.W. Oahu BISH 1928-10-01
365 Zanthoxylum kauaense Honouliuli, Kaaikukae Preserved_Specimen tree; fruit green to purple Russ, G.W. Oahu BISH 1936-10-19
366 Zanthoxylum kauaense Kilauea, Puaulu Preserved_Specimen Rock, J.F.C. 10201 Hawaii BISH 1911-07-01
367 Zanthoxylum kauaense Waimea, Kohua ridge, Kawaia trail Preserved_Specimen Cranwell, L.M. 3010 Kauai BISH 1938-08-19
368 Zanthoxylum kauaense Auwahi, S slope of Haleakala Preserved_Specimen Forbes, C.N. 1956 Maui BISH 1920-03-14
369 Zanthoxylum kauaense Kilauea, Puaulu Preserved_Specimen Rock, J.F.C. 10203 Hawaii BISH 1911-07-01
370 Zanthoxylum kauaense Honouliuli, Palawai Gulch Preserved_Specimen tree Russ, G.W. Oahu BISH 1936-10-19
371 Zanthoxylum kauaense Kamalo Gulch, above the falls Preserved_Specimen Forbes, C.N. 167 Molokai BISH 1912-07-01
372 Zanthoxylum kauaense Pohakuloa Preserved_Specimen Horner Hawaii BISH 1920-01-01
373 Zanthoxylum kauaense Waimea Canyon Preserved_Specimen Christensen, C. 194 Kauai BISH 1977-04-27
374 Zanthoxylum kauaense Kokee, Honopu Trail Preserved_Specimen small sparingly branched tree; many seedlings; leaves with 3-9 leaflets Herbst, D.R. 2158 Kauai BISH 1971-09-22
375 Zanthoxylum kauaense Honopu Trail Preserved_Specimen Leaves light green, not healthy. Leaflets 5-7 . Near Z. dipetalum (KM 39) 3 m. tall x 3 cm. DBH. Marr, K.L. 49 Kauai BISH 1988-06-11
376 Zanthoxylum kauaense Kalalau Trail Preserved_Specimen Forbes, C.N. 1091 Kauai BISH 1917-07-03
377 Zanthoxylum kauaense Waimea, Kohua Ridge, Kawaia Trail Preserved_Specimen Male plant. Cranwell, L.M. 3855 Kauai BISH 1938-08-19
378 Zanthoxylum kauaense Hapapa interval, subcrest forest of main divide Preserved_Specimen small tree 8'-9' height, only fertile ind. among several in dense layer of Syzygium and Pelea, all plants with similar foliage common in arewa,sp. distributes from ridgeliness to gulch floors Takeuchi, W.N. 3905 Oahu BISH 1988-07-17
379 Zanthoxylum kauaense Kamalo Gulch, above the falls Preserved_Specimen Forbes, C.N. 167 Molokai BISH 1912-07-01
380 Zanthoxylum kauaense Puuaulu Preserved_Specimen Forbes, C.N. 1031 Hawaii BISH 1915-07-01
381 Zanthoxylum kauaense Kaholuamanu Preserved_Specimen Rock, J.F.C. 2103 Kauai BISH 1909-03-03
382 Zanthoxylum kauaense W Waimea drainage basin Preserved_Specimen Forbes, C.N. 1159 Kauai BISH 1917-07-03
383 Zanthoxylum kauaense above Waimea Preserved_Specimen Rock, J.F.C. 5677 Kauai BISH 1909-09-01
384 Zanthoxylum kauaense Kaohai, Waiakeakua Preserved_Specimen Munro, G.C. 239 Lanai BISH 1915-04-23
385 Zanthoxylum kauaense Mahana Preserved_Specimen Rock, J.F.C. 8112 Lanai BISH 1910-08-03
386 Zanthoxylum kauaense W Waimea drainage basin Preserved_Specimen Forbes, C.N. 826 Kauai BISH 1917-07-03
387 Zanthoxylum kauaense Auwahi Preserved_Specimen Munro, G.C. 161 Maui BISH 1915-02-12
388 Zanthoxylum kauaense Mahana Preserved_Specimen Rock, J.F.C. 8112 Lanai BISH 1910-08-03
389 Zanthoxylum kauaense Schofield, Wahiwa, Waianae Mts Preserved_Specimen Warshauer, F.R. 1072 Oahu BISH 1977-02-21
390 Zanthoxylum kauaense Mts. near Koele Preserved_Specimen Forbes, C.N. 37 Lanai BISH 1913-06-01
391 Zanthoxylum kauaense Kipukaki, near Kilauea Preserved_Specimen Forbes, C.N. 403 Hawaii BISH 1911-08-04
392 Zanthoxylum kauaense Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 13046 Hawaii BISH 1917-08-01
393 Zanthoxylum kauaense Haleakala, Makawao, Waihou Gulch Preserved_Specimen Rock, J.F.C. 10200 Maui BISH 1912-03-01
394 Zanthoxylum kauaense lateral ridge, R side of Olowalu Valley Preserved_Specimen Forbes, C.N. 2255 Maui BISH 1920-05-07
395 Zanthoxylum kauaense Mahana Preserved_Specimen Rock, J.F.C. 8071 Lanai BISH 1910-06-21
396 Zanthoxylum kauaense Koolau Range, summit above Kahuku Preserved_Specimen Rock, J.F.C. 25765 Oahu BISH 1957-04-03
397 Zanthoxylum kauaense Kaholuamanu Preserved_Specimen Forbes, C.N. 320 Kauai BISH 1909-09-01
398 Zanthoxylum kauaense Waimea Canyon, Puu Ka Pele Preserved_Specimen Hobdy, R.W. 41 Kauai BISH 1968-12-01
399 Zanthoxylum kauaense Kamalo Gulch, above the falls Preserved_Specimen Forbes, C.N. 167 Molokai BISH 1912-07-01
400 Zanthoxylum kauaense Waianae Mts Preserved_Specimen Rock, J.F.C. 17054 Oahu BISH 1918-04-01
401 Zanthoxylum kauaense Kilauea, Puaulu Preserved_Specimen Rock, J.F.C. 10201 Hawaii BISH 1911-07-01
402 Zanthoxylum kauaense Mt. Kaala, Waianae Range Preserved_Specimen Gillett, G.W. 1720 Oahu BISH 1965-03-12
403 Zanthoxylum kauaense Hookio Ridge Preserved_Specimen Munro, G.C. 344 Lanai BISH 1928-06-20
404 Zanthoxylum kauaense Puuwaawaa hill Preserved_Specimen Rock, J.F.C. 3716 Hawaii BISH 1909-06-15
405 Zanthoxylum kauaense Naihakua, near Preserved_Specimen Munro, G.C. 423 Lanai BISH 1925-04-28
406 Zanthoxylum kauaense Haleakala, Makawao, Waihou Gulch Preserved_Specimen Rock, J.F.C. 10200 Maui BISH 1912-03-01
407 Zanthoxylum kauaense Waianae Mts., Dupont Trail to Puu Kaala Preserved_Specimen Krajina, V.J. 620225340 Oahu BISH 1962-02-25
408 Zanthoxylum kauaense Waianae Mts., W side of main ridge separating Makaha and Waianae Kai Valleys, N of Puu Kawiwi Preserved_Specimen Herbst, D.R. 5044 Oahu BISH 1974-09-21
409 Zanthoxylum kauaense Kilauea, Puaulu Preserved_Specimen Rock, J.F.C. 10201 Hawaii BISH 1911-07-01
410 Zanthoxylum kauaense Waianae slope of Kaala above Makaleha Preserved_Specimen Skottsberg, C.J.F. 376 Oahu BISH 1922-08-30
411 Zanthoxylum kauaense Kipuka near kilauea Preserved_Specimen Forbes, C.N. 403 Hawaii BISH 1911-12-04
412 Zanthoxylum kauaense Aukuu Preserved_Specimen Munro, G.C. 439 Lanai BISH 1915-05-29
413 Zanthoxylum kauaense Waimea Preserved_Specimen Knudsen 146 Kauai BISH
414 Zanthoxylum kauaense Lanai Preserved_Specimen Munro, G.C. 439 Lanai BISH 1915-04-29
415 Zanthoxylum kauaense Haleakala, Makawao, Waihou Gulch Preserved_Specimen Rock, J.F.C. 10200 Maui BISH 1912-03-01
416 Zanthoxylum kauaense Puuwaawaa Preserved_Specimen Rock, J.F.C. 3716 Hawaii BISH 1909-06-15
417 Zanthoxylum kauaense Olowalu Valley Preserved_Specimen Munro, G.C. 2256 Maui BISH 1920-05-07
418 Zanthoxylum kauaense Papoaea, N slope of Haleakala Preserved_Specimen Forbes, C.N. 2519 Maui BISH 1920-06-14
419 Zanthoxylum kauaense S slope of Paliloi Forest Preserved_Specimen Forbes, C.N. 2068 Maui BISH 1920-03-28
420 Zanthoxylum kauaense Hawaii Preserved_Specimen Rémy, J. 614 Hawaii BISH 1851-01-01
421 Zanthoxylum kauaense W Waimea drainage basin Preserved_Specimen Forbes, C.N. 871 Kauai BISH 1917-07-03
422 Zanthoxylum kauaense Waimea-Malaekahana, summit trail Preserved_Specimen Ozaki, E.T. 1380 Oahu BISH 1955-01-09
423 Zanthoxylum kauaense Kula, S of sanitorium Preserved_Specimen Forbes, C.N. 2176 Maui BISH 1920-04-08
424 Zanthoxylum kauaense Makawao, Haleakala Preserved_Specimen Rock, J.F.C. 10207 Maui BISH 1912-03-01
425 Zanthoxylum kauaense Waimea-Malaekahana, summit trail Preserved_Specimen Ozaki, E.T. 1380 Oahu BISH 1955-01-09
426 Zanthoxylum kauaense Kumuwela Preserved_Specimen Rice, R.C.A. 28096 Kauai BISH 1976-07-16
427 Zanthoxylum kauaense Mahana Preserved_Specimen Rock, J.F.C. 8112 Lanai BISH 1910-08-03
428 Zanthoxylum kauaense Pupukea Military Trail Preserved_Specimen MacDaniels, L.H. 544 Oahu BISH 1927-01-29
429 Zanthoxylum kauaense Kamalo Gulch, above the falls Preserved_Specimen Forbes, C.N. 167 Molokai BISH 1912-07-01
430 Zanthoxylum kauaense Waimea Canyon rim Preserved_Specimen Hobdy, R.W. 42 Kauai BISH 1968-12-09
431 Zanthoxylum kauaense Mts. near Koele Preserved_Specimen Forbes, C.N. 37 Lanai BISH 1913-06-01
432 Zanthoxylum kauaense Kaholuamanu Preserved_Specimen Rock, J.F.C. 5677 Kauai BISH 1909-09-01
433 Zanthoxylum kauaense Puaulu Kilauea volcanoes Preserved_Specimen Rock, J.F.C. 10203 Hawaii BISH 1911-07-01
434 Zanthoxylum kauaense Mountains near Koele Preserved_Specimen Forbes, C.N. 113 Lanai BISH 1913-06-01
435 Zanthoxylum kauaense Kawela Valley Preserved_Specimen Cranwell, L.M. 3526 Molokai BISH 1938-06-26
436 Zanthoxylum kauaense Mahana Preserved_Specimen Rock, J.F.C. 8112 Lanai BISH 1910-08-03
437 Zanthoxylum kauaense East Maui, West Kaupo in kipuka in a Aa lava flow. Preserved_Specimen Welton, P. 1999 Maui BISH 1997-10-01
438 Zanthoxylum kauaense Kanehoa ridge, 300' below summit Preserved_Specimen Pyle, L. 2029 Oahu BISH 1985-03-03
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